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beach sand-spurrey, beach sand-spurry, sticky sand spurry, sticky sand-spurrey

bristle-seed sand-spurrey, bristleseed sandspurry

Habit Plants strongly perennial with branched, woody base, stout, 5–40 cm, densely stipitate-glandular in inflorescence or throughout. Plants annual, delicate, 5–15 cm, stipitate-glandular throughout.
Taproots

becoming stout, woody.

slender.

Stems

erect or ascending to prostrate, usually branched proximally;

main stem 0.8–3 mm diam. proximally.

erect to ascending, simple to diffusely branched proximally and distally;

main stem occasionally prostrate, 0.2–1.5 mm diam. proximally.

Leaves

stipules conspicuous, dull white to tan, narrowly triangular, 4.5–11 mm, apex long-acuminate;

blade linear, (0.6–)1–5.5 cm, fleshy, apex apiculate to spine-tipped;

axillary leaves 1–2+ per cluster.

stipules inconspicuous, silvery to dull tan, broadly triangular, 1.4–2.4 mm, shorter than wide, apex acuminate;

blade linear, 0.5–3.5 cm, somewhat fleshy, apex blunt to apiculate;

axillary leaf cluster usually absent.

Pedicels

erect, divergent, or reflexed in fruit.

reflexed and oriented to 1 side in fruit.

Flowers

sepals connate 0.5–1.8 mm proximally, lobes often 3-veined, ovate to lanceolate, 4.5–7 mm, to 8 mm in fruit, margins 0.3–0.7 mm wide, apex blunt to rounded;

petals white or pink to rosy, elliptic to obovate, 0.9–1.3 times as long as sepals in flower;

stamens 9–10;

styles 0.6–3 mm.

sepals connate 0.2–0.3 mm proximally, lobes 1–3-veined, lanceolate to ovate, 2.5–3.6 mm, to 4 mm in fruit, margins 0.1–0.5 mm wide, apex rounded;

petals white to pink or rosy, lanceolate, 0.4–0.6 times as long as sepals;

stamens 1–4(–5);

styles 0.3–0.4 mm.

Capsules

tan, 4.6–10 mm, 0.8–1.4 times as long as sepals.

greenish to tan, 2.8–4 mm, 0.9–1.4 times as long as sepals.

Seeds

± red-brown, often with submarginal groove or depression, suborbiculate to pyriform, compressed, (0.6–)0.7–1.2 mm, smooth to faintly or prominently tuberculate or sculpturing of parallel, wavy lines or of low, rounded mounds, not papillate;

wing absent or white to reddish brown proximally, 0.1–0.3 mm wide, margins irregular.

reddish brown to blackish, silver tinged, with submarginal groove, pyriform, ± compressed, 0.5–0.7(–0.8) mm, shiny, slightly roughened, with dense, gland-tipped papillae and appearing echinate (30x);

wing usually absent, whitish to reddish brown, 0.1–0.2 mm wide, margins irregular and not papillate.

Cymes

simple to 3-compound or flowers solitary and axillary.

commonly 4–8+-compound.

2n

= 18, 36 (both Europe).

Spergularia macrotheca

Spergularia echinosperma

Phenology Flowering spring.
Habitat Dunes, clay flats, sandy river banks
Elevation 0-20+ m (0-100+ ft)
Distribution
from FNA
CA; OR; WA; BC; including nw Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; GA; LA; TX; Europe [Introduced in North America]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals pink to rosy; styles 0.6-1.2 mm
var. macrotheca
1. Petals white; styles 1.2-3 mm
→ 2
2. Capsules 1.2-1.4 times as long as sepals; styles 1.2-1.9 mm
var. leucantha
2. Capsules 0.8-1 times as long as sepals; styles 2-3 mm
var. longistyla
Source FNA vol. 5, p. 18. FNA vol. 5, p. 22.
Parent taxa Caryophyllaceae > subfam. Polycarpoideae > Spergularia Caryophyllaceae > subfam. Polycarpoideae > Spergularia
Sibling taxa
S. atrosperma, S. bocconi, S. canadensis, S. diandra, S. echinosperma, S. media, S. platensis, S. rubra, S. salina, S. villosa
S. atrosperma, S. bocconi, S. canadensis, S. diandra, S. macrotheca, S. media, S. platensis, S. rubra, S. salina, S. villosa
Subordinate taxa
S. macrotheca var. leucantha, S. macrotheca var. longistyla, S. macrotheca var. macrotheca
Synonyms Arenaria macrotheca
Name authority (Hornemann ex Chamisso & Schlechtendal) Heynhold: Alph. Aufz. Gew. 689. (1846) Celakovský: Arch. Naturwiss. Landesdurchf. Bohmen 4: 867. (1881)
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