Spergularia macrotheca |
Spergularia bocconi |
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beach sand-spurrey, beach sand-spurry, sticky sand spurry, sticky sand-spurrey |
Boccone's sand spurry, Boccone's sand-spurrey, Boccone's sea-spurrey, Bocconi's sand-spurrey |
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Habit | Plants strongly perennial with branched, woody base, stout, 5–40 cm, densely stipitate-glandular in inflorescence or throughout. | Plants annual, ± delicate, 6–25+ cm, often densely stipitate-glandular, at least in inflorescence. | ||||||||
Taproots | becoming stout, woody. |
± filiform. |
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Stems | erect or ascending to prostrate, usually branched proximally; main stem 0.8–3 mm diam. proximally. |
erect to spreading or sprawling, usually much-branched proximally; main stem 0.5–1 mm diam. proximally. |
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Leaves | stipules conspicuous, dull white to tan, narrowly triangular, 4.5–11 mm, apex long-acuminate; blade linear, (0.6–)1–5.5 cm, fleshy, apex apiculate to spine-tipped; axillary leaves 1–2+ per cluster. |
stipules usually inconspicuous, dull white to tan, broadly triangular, 1.5–4.5 mm, apex acute to short-acuminate; blade ± linear, 1–4.2 cm, at least moderately fleshy, apex apiculate to spine-tipped; axillary leaves absent or 1–2 per cluster. |
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Pedicels | erect, divergent, or reflexed in fruit. |
often oriented to 1 side in fruit. |
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Flowers | sepals connate 0.5–1.8 mm proximally, lobes often 3-veined, ovate to lanceolate, 4.5–7 mm, to 8 mm in fruit, margins 0.3–0.7 mm wide, apex blunt to rounded; petals white or pink to rosy, elliptic to obovate, 0.9–1.3 times as long as sepals in flower; stamens 9–10; styles 0.6–3 mm. |
sepals connate 0.3–0.6 mm proximally, lobes often 3-veined, ovate to elliptic-oblong, 2.2–3.5 mm, to 4.5 mm in fruit, margins 0.2–0.5 mm wide, apex acute to rounded; petals white or pink to rosy, ovate to obovate, 0.8–1 times as long as sepals; stamens 8–10; styles 0.4–0.6 mm. |
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Capsules | tan, 4.6–10 mm, 0.8–1.4 times as long as sepals. |
greenish, 3(–4) mm, 1–1.2 times as long as sepals. |
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Seeds | ± red-brown, often with submarginal groove or depression, suborbiculate to pyriform, compressed, (0.6–)0.7–1.2 mm, smooth to faintly or prominently tuberculate or sculpturing of parallel, wavy lines or of low, rounded mounds, not papillate; wing absent or white to reddish brown proximally, 0.1–0.3 mm wide, margins irregular. |
light brown, with submarginal groove, broadly ovate, plump, 0.4–0.6 mm, somewhat shiny, smooth to minutely roughened, margins with peglike papillae (40x); wing absent. |
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Cymes | simple to 3-compound or flowers solitary and axillary. |
simple to 6+-compound. |
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2n | = 18? (Africa), 36 (Europe). |
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Spergularia macrotheca |
Spergularia bocconi |
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Phenology | Flowering spring. | |||||||||
Habitat | Salt marshes, alkaline places, sandy soils | |||||||||
Elevation | 0-400 m (0-1300 ft) | |||||||||
Distribution |
CA; OR; WA; BC; including nw Mexico
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CA; OR; sw Europe (Mediterranean region) [Introduced in North America]
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Discussion | Varieties 3 (3 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The spelling of the epithet bocconi, often “corrected” to bocconii, is debatable. It commemorates Paolo Boccone, suggesting a correction to bocconei, but he also used the Latinized form Bocconus, allowing bocconi. We have used bocconi, following the first usage by Scheele. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 5, p. 18. | FNA vol. 5, p. 20. | ||||||||
Parent taxa | Caryophyllaceae > subfam. Polycarpoideae > Spergularia | Caryophyllaceae > subfam. Polycarpoideae > Spergularia | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Arenaria macrotheca | Alsine bocconi | ||||||||
Name authority | (Hornemann ex Chamisso & Schlechtendal) Heynhold: Alph. Aufz. Gew. 689. (1846) | (Scheele) Graebner: in P. F. A. Ascherson et al., Syn. Mitteleur. Fl. 5(1): 849. (1919) | ||||||||
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