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beach sand-spurrey, beach sand-spurry, sticky sand spurry, sticky sand-spurrey

black-seed sand-spurrey, blackseed sandspurry, sand spurrey

Habit Plants strongly perennial with branched, woody base, stout, 5–40 cm, densely stipitate-glandular in inflorescence or throughout. Plants annual, delicate, 6–15 cm, glabrous or stipitate-glandular distally.
Taproots

becoming stout, woody.

filiform.

Stems

erect or ascending to prostrate, usually branched proximally;

main stem 0.8–3 mm diam. proximally.

erect to ascending or arcuate-spreading, simple or branched proximally;

main stem 0.3–1 mm diam. proximally.

Leaves

stipules conspicuous, dull white to tan, narrowly triangular, 4.5–11 mm, apex long-acuminate;

blade linear, (0.6–)1–5.5 cm, fleshy, apex apiculate to spine-tipped;

axillary leaves 1–2+ per cluster.

stipules inconspicuous, dull white to tan, broadly triangular, 1–2.5 mm, apex acute to short-acuminate;

blade filiform to linear, 0.5–3.5 cm, fleshy, apex blunt to apiculate;

axillary leaf clusters absent.

Pedicels

erect, divergent, or reflexed in fruit.

divergently spreading to reflexed in fruit.

Flowers

sepals connate 0.5–1.8 mm proximally, lobes often 3-veined, ovate to lanceolate, 4.5–7 mm, to 8 mm in fruit, margins 0.3–0.7 mm wide, apex blunt to rounded;

petals white or pink to rosy, elliptic to obovate, 0.9–1.3 times as long as sepals in flower;

stamens 9–10;

styles 0.6–3 mm.

sepals connate 0.2–0.5 mm proximally, lobes ± weakly 1-veined, elliptic to ovate, 1.8–2.7 mm, to 3.5 mm in fruit, margins 0.1–0.5 mm wide, apex broadly acute to rounded;

petals white to rosy, ± ovate, 0.7–0.8 times as long as sepals;

stamens 4–8;

styles 0.4–0.8 mm.

Capsules

tan, 4.6–10 mm, 0.8–1.4 times as long as sepals.

greenish, 3.3–5 mm, 1–1.3 times as long as sepals.

Seeds

± red-brown, often with submarginal groove or depression, suborbiculate to pyriform, compressed, (0.6–)0.7–1.2 mm, smooth to faintly or prominently tuberculate or sculpturing of parallel, wavy lines or of low, rounded mounds, not papillate;

wing absent or white to reddish brown proximally, 0.1–0.3 mm wide, margins irregular.

black, often iridescent, submarginal groove usually absent, broadly ovate, compressed, 0.6–0.8 mm, shiny, sculpturing of parallel, wavy lines, not papillate (40x);

wing absent or, rarely, incomplete, brownish, 0.1–0.2 mm, margins irregular.

Cymes

simple to 3-compound or flowers solitary and axillary.

simple to 2-compound.

Spergularia macrotheca

Spergularia atrosperma

Phenology Flowering summer.
Habitat Alkaline places, mud flats, stream beds, sandy places
Elevation 30-1500 m (100-4900 ft)
Distribution
from FNA
CA; OR; WA; BC; including nw Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV
Discussion

Varieties 3 (3 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petals pink to rosy; styles 0.6-1.2 mm
var. macrotheca
1. Petals white; styles 1.2-3 mm
→ 2
2. Capsules 1.2-1.4 times as long as sepals; styles 1.2-1.9 mm
var. leucantha
2. Capsules 0.8-1 times as long as sepals; styles 2-3 mm
var. longistyla
Source FNA vol. 5, p. 18. FNA vol. 5, p. 19.
Parent taxa Caryophyllaceae > subfam. Polycarpoideae > Spergularia Caryophyllaceae > subfam. Polycarpoideae > Spergularia
Sibling taxa
S. atrosperma, S. bocconi, S. canadensis, S. diandra, S. echinosperma, S. media, S. platensis, S. rubra, S. salina, S. villosa
S. bocconi, S. canadensis, S. diandra, S. echinosperma, S. macrotheca, S. media, S. platensis, S. rubra, S. salina, S. villosa
Subordinate taxa
S. macrotheca var. leucantha, S. macrotheca var. longistyla, S. macrotheca var. macrotheca
Synonyms Arenaria macrotheca
Name authority (Hornemann ex Chamisso & Schlechtendal) Heynhold: Alph. Aufz. Gew. 689. (1846) R. Rossbach: Rhodora 42: 80, plate 589, fig.3a-c.1940
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