Spergula arvensis |
Caryophyllaceae subfam. polycarpoideae |
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corn-spurrey, corn-spurry, spargoute des champs, starwort, stickwort |
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Habit | Plants glabrous or, often, glandular. | Herbs [small shrubs], winter annual, annual, or perennial; taprooted, not rhizomatous. |
Stems | usually branched proximally, 10–50+ cm. |
prostrate to erect, simple or branched. |
Leaves | blades usually appearing terete, 1.5–3(–5) cm, margins often revolute, forming abaxial channel. |
opposite or sometimes appearing whorled, bases connate or not, sometimes petiolate or often sessile, stipulate; stipules ovate or deltate to lanceolate or bristlelike, scarious; blade subulate or subtriangular to linear and threadlike or spatulate to ovate or orbiculate, seldom succulent. |
Inflorescences | terminal or axillary cymes, or flowers solitary; bracts scarious or absent; involucel bracteoles absent. |
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Pedicels | erect to ascending, reflexed, secund in fruit. |
present or flowers sessile. |
Flowers | sepals 3.5–5 mm; petals ovate, 3/4–1 times as long as sepals in flower, apex obtuse; stamens usually 10. |
bisexual or rarely unisexual; perianth and androecium hypogynous or perigynous; hypanthium absent or dish- or cup-shaped; sepals 5, distinct or sometimes connate proximally, hooded (Drymaria, Polycarpon) or not, awned (Polycarpon) or not; petals absent or (3–)5, blade clawed (Drymaria) or not, auricles absent, coronal appendages absent, blade apex entire, erose, or 2(–4)-fid, sometimes emarginate; stamens (1–)3–5(–10), in 1 or 2 whorls, usually arising from base of ovary or from rim of hypanthium (Spergula, Spergularia); staminodes absent; ovary 1-locular; styles 1 or 3, occasionally 2 or 5, distinct or sometimes connate proximally; stigmas 3, occasionally 2 or 5. |
Fruits | capsules, opening by 3 or 5, occasionally 4 valves; carpophore sometimes present. |
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Capsule | valves 3.5–5 mm. |
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Seeds | sometimes keeled or winged, subglobose, 1–1.1 mm wide, surface minutely roughened or obscurely low-tuberculate (50x), covered with white, club-shaped papillae in part or throughout (packing of seeds in capsule may prevent papillae development in spots), wings white, ± 0.1 mm wide. |
3–150+, whitish or tan to often brown or black, ± triangular, pyriform, or reniform to circular, subglobose or laterally compressed to angular (Polycarpon); embryo peripheral, curved or rarely annular to spirally curved (Spergula). |
x | = [7], 8, 9, (11), 12. |
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2n | = 18, 36 (both Europe). |
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Spergula arvensis |
Caryophyllaceae subfam. polycarpoideae |
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Phenology | Flowering spring–early summer. | |
Habitat | Sandy roadsides, cultivated fields, other disturbed areas | |
Elevation | 10-2000 m (0-6600 ft) | |
Distribution |
AK; AL; AR; CA; CO; CT; DC; DE; FL; GA; ID; IL; IN; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; NH; NJ; NY; OH; OR; PA; RI; SC; TX; VA; VT; WA; WI; WV; WY; AB; BC; NB; NF; NS; NT; ON; PE; QC; SK; YT; SPM; Greenland; Eurasia [Introduced in North America; introduced in Central America, South America, Asia (Korea), Africa, Australia]
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w North America (including Mexico); w South America; Europe (Mediterranean region); Asia (Mediterranean region); Africa (Mediterranean and tropical regions); less diverse in temperate areas |
Discussion | Spergula arvensis is often a significant weed in sandy crop lands, but it is sometimes used as a forage crop in areas with poor, sandy soils; it was intentionally introduced to Crawford County, Michigan, in 1888 (O. Clute and O. Palmer 1893). Historical collections are known also from Kansas, Nebraska, North Dakota, and South Dakota, where Spergula arvensis may have been introduced but apparently did not persist. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 16, species ca. 210 (7 genera, 28 species in the flora). More commonly recognized as a tribe, Polycarpoideae is characterized by the presence of stipules, well-developed petals, and capsules. It is a relatively small group, with about four-fifths of the species in Drymaria, Polycarpaea, and Spergularia. Although clearly stipulate, Spergularia clusters with members of Alsinoideae in recent molecular studies (M. Nepokroeff et al. 2002; R. D. Smissen et al. 2002). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 5, p. 15. | FNA vol. 5, p. 9. |
Parent taxa | Caryophyllaceae > subfam. Polycarpoideae > Spergula | Caryophyllaceae |
Sibling taxa | ||
Subordinate taxa | ||
Synonyms | S. arvensis var. sativa | |
Name authority | Linnaeus: Sp. Pl. 1: 440. 1753 | Tanfani: in F. Parlatore, Fl. Ital. 9: 623. (1892) |
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