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California cord grass, Pacific cordgrass

Chilean cordgrass, dense-flower cord grass, densely-flower cordgrass

Habit Plants occasionally streaked or tinged with purple, rhizomatous; rhizomes elongate, flaccid, whitish, scales inflated, not closely imbricate. Plants cespitose, rarely rhizomatous; rhizomes, when present, short, to 10 mm thick.
Culms

to 150 cm tall, to 10 mm thick, erect, terete, solitary or in small clumps, succulent, glabrous, often with adventitious roots from the lower nodes, having an unpleasant, sulphurous odor when fresh.

27-150 cm, forming large clumps, indurate, usually with short extravaginal shoots appressed to the culms.

Sheaths

mostly glabrous, throats sparsely pilose, lower sheaths sometimes somewhat wrinkled;

ligules 1-2 mm;

blades 8-12 mm wide, flat to loosely involute, glabrous, margins usually smooth, sometimes slightly scabrous, apices acuminate.

glabrous, lower sheaths smooth, indurate and shining, upper sheaths dull and somewhat striate;

ligules 1-2 mm;

blades 12-43 cm long, 3-8 mm wide, involute when fresh, abaxial surfaces glabrous, adaxial surfaces and margins scabrous, apices acuminate.

Panicles

12-25 cm, with 3-25 branches, smoothly cylindrical, often partially enclosed in the uppermost sheath;

rachises twisted, glabrous;

branches 2-8 cm, usually closely appressed and twisted, lower branches noticeably longer and less closely imbricate than the upper branches, all branches with axes rarely extending past the distal spikelets, with 8-30 spikelets.

10-30 cm long, 4-8 mm wide, sinuous in outline, often twisted, with 2-15 branches;

branches 1-11 cm long, longer branches narrower than the shorter branches, all branches tightly appressed, moderately imbricate, axes not prolonged beyond the distal spikelets, with 10-30 spikelets.

Spikelets

8-25 mm, usually appressed, often appearing twisted, those on the lower branches usually less closely imbricate than those on the upper branches.

8-14 mm.

Glumes

usually curved, sides and keels glabrous, scabrous, or hispid, apices acuminate to obtuse or rounded;

lower glumes 6-12 mm;

upper glumes 8-25 mm, 1-veined;

lemmas glabrous or sparsely appressed pubescent on the sides, keels glabrous, apices obtuse, rounded or lobed;

paleas slightly exceeding the lemmas, thin, papery, glabrous, apices usually rounded, rarely acuminate;

anthers 3-6 mm.

glabrous or sparsely hispidulous, keels hispidulous, margins sparsely hispidulous;

lower glumes 4-7 mm, usually obtuse;

upper glumes 8-14 mm, 1-veined, usually acuminate;

lemmas minutely hispidulous, keels glabrous proximally, hispidulous distally, apices acuminate to obtuse;

paleas acuminate, keels glabrous basally, hispidulous distally;

anthers 3-5 mm.

2n

= 60.

= 60 [this chromosome count was obtained by Gerish (1979), who reported it for S. foliosa, but Spicher and Josselyn (1985) demonstrated that the plants he worked with were almost certainly S. densiflora, a species that hitherto had been misidentified as the native S. foliosa].

Spartina foliosa

Spartina densiflora

Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR; WA; BC
[BONAP county map]
Discussion

Spartina foliosa grows in the intertidal zone from northern California to Baja California, Mexico. Populations in San Francisco Bay are threatened by various introduced species of Spartina. Of particular concern is S. alterniflora, which forms hybrids with S. foliosa that have a broader ecological amplitude than either parent. In California, S. foliosa is often confused with S. densiflora, which is also established in some regions, but S. foliosa differs from that species in being rhizomatous and having softer culms and wider leaf blades.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Spartina densiflora is native to South America, where it grows in coastal marshes and at inland sites. It was introduced to Humboldt Bay, Humboldt County, California, possibly during the nineteenth century. It is now established there and in several locations around San Francisco Bay and in Washington, Oregon, and Texas, as well as the Mediterranean coast of Europe. In California, it has often been mistaken for S. foliosa, from which it differs in its indurate culms, narrow, inrolled leaves, and cespitose growth habit and tendency to grow among Salicornia in the upper intertidal zone or in open mud.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 25, p. 244. FNA vol. 25, p. 247.
Parent taxa Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Spartina Poaceae > subfam. Chloridoideae > tribe Cynodonteae > Spartina
Sibling taxa
S. alterniflora, S. anglica, S. bakeri, S. cynosuroides, S. densiflora, S. gracilis, S. maritima, S. patens, S. pectinata, S. spartinae, S. ×caespitosa, S. ×townsendii
S. alterniflora, S. anglica, S. bakeri, S. cynosuroides, S. foliosa, S. gracilis, S. maritima, S. patens, S. pectinata, S. spartinae, S. ×caespitosa, S. ×townsendii
Name authority Trin. Brongn.
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