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California Mountain Ashe, California mountain-ash

Sitka mountain-ash, western mountain ash

Habit Shrubs, 10–20(–40) dm. Shrubs, 10–40 dm.
Stems

1–8;

bark dark gray;

winter buds red to brown, shiny, conic, 5–12 mm, slightly glutinous, sparsely hairy except along scale margins and apex, hairs rufous.

1–8;

bark initially reddish purple to brownish gray, becoming grayish red;

winter buds red-purple to red-brown, conic to ovoid, 8–13 mm, dull, slightly glaucous, not glutinous, sparsely or densely villous, hairs primarily rufous.

Leaves

pinnately compound;

stipules often persistent, margins rufous-hairy;

blade pale abaxially, shiny, green adaxially, leaflets 7–9(–11), opposite or subopposite, oblong to elliptic, sometimes narrowly elliptic, 2.5–4.1(–4.5) × 0.9–2 cm, l/w ratio 2.1–2.6, margins finely to coarsely serrate, sometimes doubly serrate, apex obtuse to acute, surfaces soon glabrous, leaflet axils and petiole bases hairy, hairs rufous.

pinnately compound;

stipules persistent, sometimes deciduous, hairs rufous;

blade paler abaxially, dull blue-green, usually slightly glaucous, rarely faintly shiny adaxially, leaflets 7–13, opposite, sometimes subopposite, oblong to narrowly elliptic, sometimes oblanceolate, ovate, or obovate, 1.8–6 × 0.6–2.5 cm, l/w ratio 1.9–3.5, margins entire or finely to coarsely serrate, sometimes doubly serrate, apex obtuse or acute, surfaces soon glabrous, leaf and leaflet axils hairy, hairs rufous.

Panicles

25–120+-flowered, rounded, 3–11 cm diam.;

peduncles glabrous or sparsely hairy.

12–80-flowered, rounded, 2–8 cm diam.;

peduncles glabrous, glaucous, or sparsely to densely villous, hairs primarily rufous.

Pedicels

glabrous or sparsely hairy; fruiting pedicels essentially glabrous.

glabrous, glaucous, or sparsely to densely villous, hairs primarily rufous.

Flowers

9–10 mm diam.;

hypanthium glabrous, hypanthium plus sepals 3 mm;

sepals 0.7–1.5 mm, margins ciliolate, hairs whitish, irregularly glandular;

petals white, broadly ovate, 3–4 mm;

stamens 20;

carpels distinct, apex conic, styles 3 or 4, 1.5–2 mm.

11–17 mm diam.;

hypanthium glabrous, glaucous, hypanthium plus sepals 3–4 mm;

sepals 1–2.2 mm, margins lightly to moderately ciliate, hairs rufous or whitish, infrequently glandular;

petals white, rarely pinkish, rhombic to ovate, 4–7.5 mm;

stamens 15–20;

carpels distinct, apex concave, depressed, or slightly conic and truncate, styles 3–5, 1.5–3.2 mm.

Infructescences

glabrous or nearly so.

glabrous, glaucous, or villous.

Pomes

bright red, globose to subglobose, 6–9 mm diam., shiny if glaucous layer is rubbed off;

sepals inconspicuous, incurved.

pinkish red or red (often appearing slightly purplish), subglobose to broadly elliptic or obovoid, 7–13 × 7–13 mm, dull, glaucous;

sepals inconspicuous, incurved.

Seeds

red-brown, lanceolate, 4 × 2 mm, slightly asymmetric, slightly flattened.

red-brown, ovoid to ovoid-lanceolate, oblong or elliptic, 3.4–5 × 2.2–2.9 mm, symmetric or trigonous, slightly flattened.

Sorbus californica

Sorbus sitchensis

Phenology Flowering spring; fruiting fall.
Habitat Mountain slopes, meadows, stream banks, lakeshores, sun or shade
Elevation 1500–3400 m (4900–11200 ft)
Distribution
from FNA
CA; NV; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; CA; ID; MT; OR; WA; AB; BC; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Sorbus californica is recognized here in the narrow sense, occurring north to Crater Lake National Park, Oregon (P. F. Zika 2003). The concept in H. A. McAllister (2005) differs, including plants of British Columbia and Washington treated here as S. scopulina. Sorbus californica is distinct in its shiny leaflets less than 4 cm with rufous hairs in the axils. Where their ranges overlap, S. scopulina has hairier inflorescences, leaflets usually more than 4.5 cm, and whitish axillary hairs. The small leaflets and essentially glabrous inflorescences help distinguish S. californica from putative hybrids between S. scopulina and S. sitchensis found farther north, which are otherwise similar with their somewhat shiny leaflets, slightly glaucous fruits, and red axillary hairs. Some collections of S. californica from California are intermediate with either S. sitchensis or S. scopulina and may represent hybrids with those two species.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

The other western Sorbus with rufous indument in the leaf and leaflet axils is S. californica, which differs in its smaller, shinier leaflets, less hairy inflorescences, and shiny winter buds. The winter buds of S. sitchensis are hairier and less scaly at the summit, also glaucous and never shiny or glutinous. In the field, the fresh fruits of S. sitchensis are often pinkish red; fruits of S. californica and S. scopulina are generally orange-red.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaflet margins toothed apically, serrate for more than 1/2 their length; petals white.
var. sitchensis
1. Leaflet margins entire or toothed apically, serrate for less than 1/2 their length; petals white or pinkish.
var. grayi
Source FNA vol. 9, p. 441. FNA vol. 9, p. 444.
Parent taxa Rosaceae > subfam. Amygdaloideae > tribe Maleae > Sorbus > subg. Sorbus > sect. Commixtae Rosaceae > subfam. Amygdaloideae > tribe Maleae > Sorbus > subg. Sorbus > sect. Tianshanicae
Sibling taxa
S. americana, S. aucuparia, S. decora, S. hybrida, S. intermedia, S. sambucifolia, S. scopulina, S. sitchensis, S. torminalis
S. americana, S. aucuparia, S. californica, S. decora, S. hybrida, S. intermedia, S. sambucifolia, S. scopulina, S. torminalis
Subordinate taxa
S. sitchensis var. grayi, S. sitchensis var. sitchensis
Synonyms S. sitchensis subsp. californica
Name authority Greene: Pittonia 4: 131. (1900) M. Roemer: Fam. Nat. Syn. Monogr. 3: 139. (1847)
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