Sorbus californica
Sorbus decora
California Mountain Ashe, California mountain-ash
northern mountain-ash, showy mountain-ash, sorbier plaisant
1–8;
bark dark gray;
winter buds red to brown, shiny, conic, 5–12 mm, slightly glutinous, sparsely hairy except along scale margins and apex, hairs rufous.
1–4+;
bark gray to bronze;
winter buds purplish at maturity, shiny, conic, 10–20 mm, glutinous, scales ciliate to moderately (densely) villous, hairs primarily rufous.
pinnately compound;
stipules often persistent, margins rufous-hairy;
blade pale abaxially, shiny, green adaxially, leaflets 7–9(–11), opposite or subopposite, oblong to elliptic, sometimes narrowly elliptic, 2.5–4.1(–4.5) × 0.9–2 cm, l/w ratio 2.1–2.6, margins finely to coarsely serrate, sometimes doubly serrate, apex obtuse to acute, surfaces soon glabrous, leaflet axils and petiole bases hairy, hairs rufous.
pinnately compound;
stipules deciduous, sometimes persistent, glabrous or with whitish and/or rufous hairs;
blade paler abaxially, dull green to bluish green adaxially, leaflets (11–)13–17, opposite, elliptic, oblong to oblong-ovate, 4–7(–8) × 1.5–2.5 cm, l/w ratio 2.4–3.2(–3.6), margins serrate, sometimes serrulate, at least in distal 1/2 and often almost to base, apex abruptly short-acuminate, cuspidate, acute, or obtuse, surfaces essentially glabrous or sparsely (moderately) villous at flowering, usually glabrescent thereafter, sometimes persistently villous along midveins abaxially;
leaflet axils and petiole bases glabrous or with some rufous and/or whitish hairs adaxially.
25–120+-flowered, rounded, 3–11 cm diam.;
peduncles glabrous or sparsely hairy.
75–400+-flowered, flat-topped or rounded, 6–15 cm diam.;
peduncles sparsely to moderately (densely) villous.
glabrous or sparsely hairy; fruiting pedicels essentially glabrous.
sparsely to moderately (densely) villous.
9–10 mm diam.;
hypanthium glabrous, hypanthium plus sepals 3 mm;
sepals 0.7–1.5 mm, margins ciliolate, hairs whitish, irregularly glandular;
petals white, broadly ovate, 3–4 mm;
stamens 20;
carpels distinct, apex conic, styles 3 or 4, 1.5–2 mm.
(7–)8–12 mm diam.;
hypanthium glabrous or sparsely villous proximally, hairs whitish or rufous, hypanthium plus sepals 2.5–3.5 mm;
sepals 1–1.5(–2) mm, margins entire, often with a few thick glands;
petals white, orbiculate to obovate, (3–)3.5–5 mm;
stamens 15–20;
carpels 1/2 adnate to hypanthium, apex conic, styles 3 or 4, 1.5–2.5 mm.
glabrous or nearly so.
sparsely to moderately villous, rarely glabrous.
bright red, globose to subglobose, 6–9 mm diam., shiny if glaucous layer is rubbed off;
sepals inconspicuous, incurved.
bright red, globose to subglobose, (5–)7–11 mm diam., shiny or dull, often glaucous when dried;
sepals inconspicuous, incurved.
red-brown, lanceolate, 4 × 2 mm, slightly asymmetric, slightly flattened.
brown, lanceoloid, 4.5–5 × 1.5–2 mm, slightly asymmetric, slightly flattened.
= 68.
Sorbus californica
Sorbus decora
Sorbus californica is recognized here in the narrow sense, occurring north to Crater Lake National Park, Oregon (P. F. Zika 2003). The concept in H. A. McAllister (2005) differs, including plants of British Columbia and Washington treated here as S. scopulina. Sorbus californica is distinct in its shiny leaflets less than 4 cm with rufous hairs in the axils. Where their ranges overlap, S. scopulina has hairier inflorescences, leaflets usually more than 4.5 cm, and whitish axillary hairs. The small leaflets and essentially glabrous inflorescences help distinguish S. californica from putative hybrids between S. scopulina and S. sitchensis found farther north, which are otherwise similar with their somewhat shiny leaflets, slightly glaucous fruits, and red axillary hairs. Some collections of S. californica from California are intermediate with either S. sitchensis or S. scopulina and may represent hybrids with those two species.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Sorbus decora reputedly flowers about one week later than S. americana. The two are sometimes confused when one is not familiar with the smaller flowers and fruits, and the distinctly longer and notably acuminate leaflets of S. americana. Leaflet length/width ratios greater than 3.2 are sometimes observed on narrower leaflets of S. decora, and a range of representative leaflets need to be sampled before determination. Another species frequently confused with S. decora is S. aucuparia, characterized by ovoid, usually densely whitish-villous and not glutinous winter buds as well as densely villous flowering pedicels and hypanthia, and often abaxially tomentose leaflets at flowering. The teeth on margins of leaves in S. decora are more finely pointed and sharper than those of S. aucuparia, and the apical tooth is often distinctly prolonged in S. decora; it is no more conspicuous than the lateral ones in S. aucuparia (E. G. Voss 1972–1996, vol. 2). The leaflets of S. aucuparia are reported to be minutely papillose abaxially (visible at 100× ); those of S. decora and other members of sect. Commixtae are not papillose (H. A. McAllister 2005). Sorbus decora is reported to be extirpated from Indiana.
Chromosome counts of 2n = 34 were reported by Á. Löve and D. Löve (1965, 1982b) for Sorbus decora. Doubt exists about the identity of the material that they sampled, as the only voucher specimen retraced (Á. & D. Löve 6559, COLO, WIN), although only vegetative and juvenile, is nonetheless referable to S. aucuparia. These doubtful chromosome counts also formed the basis for the recognition of S. groenlandica (Löve and Löve 1965b), reputedly of Greenland, coastal Labrador, and alpine and subalpine regions of northeastern North America, which apparently differed from S. decora with a chromosome count of 2n = 68 (T. W. Böcher and K. Larsen 1950; K. Holmen in C. A. Jørgensen et al. 1958). Reliable S. decora chromosome counts by H. A. McAllister (2005) of three wild collections (Dundas, Ontario; St. Anne de Beaupré, Quebec; Narsarsuaq Greenland) and two of cultivated material rendered this proposition untenable with consistent counts of 2n = 68. Further, critical examination of herbarium specimens from Greenland yielded no consistent morphologic character to separate S. decora from S. groenlandica, though leaflet and fruit size in the latter tended to be relatively smaller.
A naturally occurring putative intergeneric hybrid, ×Sorbaronia arsenii (Britton ex L. Arsène) G. N. Jones, synonym Pyrus ×arsenii (Britton ex L. Arsène) L. Arsène [= Aronia ×prunifolia (Marshall) Rehder × Sorbus decora], is known from New Brunswick, Nova Scotia, Prince Edward Island, St. Pierre and Miquelon, Quebec, and, possibly, Newfoundland. The hybrid is a 10–25 dm shrub with finely serrulate, partially pinnate leaves containing (0 or)1–3 pairs of reduced, distinct leaflets proximally and a much larger, 1–3-lobed terminal segment. The midvein on the adaxial surface of the terminal leaflet may be irregularly glandular or eglandular. A possible variant of this hybrid (parentage unconfirmed), much closer to Sorbus in appearance, is found in southwestern Newfoundland. It possesses finely serrulate, almost fully pinnate leaves, often with four or five pairs of distinct leaflets, the distal first or second leaflet pairs more or less pinnatifid or partially fused to the terminal leaflet. The hypanthium and abaxial surface of the sepals are villous, the anthers are pink or red, and the pomes are usually red, though some are reported to be blackish in age, often with some tomentum persisting on the abaxial surface of the persistent sepals.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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