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shaggy hedge-hyssop

hedge-hyssop

Habit Herbs, perennial; caudex thick, hard.
Stems

erect, 20–65 cm, villous, rarely glabrous.

ascending to erect, villous (hairs jointed), rarely glabrous.

Leaves

blade lanceolate to ovate, 8–20 × 4–13 mm, base clasping, rounded to subcordate, margins entire or serrate, not or slightly revolute, apex rounded to obtuse or acute, surfaces villous, rarely glabrous, papillate, distinctly punctate abaxially.

cauline, sometimes also basal, opposite;

petiole absent, sometimes present on basal leaves;

blade not fleshy, leathery, margins entire or serrate.

Inflorescences

axillary, flowers 1 or 2 per node;

bracts present.

Pedicels

0–3(–5) mm;

bracteoles 6–9(–11) mm.

present or absent;

bracteoles present, 2.

Flowers

calyx lobes 5–7 × 0.5–1.5 mm;

corolla tubular, 5–9 mm, tube yellow, veins bluish purple, limb white.

bisexual;

sepals 5, distinct, linear to linear-lanceolate, calyx ± bilaterally symmetric, campanulate;

corolla white, sometimes tinged purple or pink, bilaterally symmetric, bilabiate to bilabiate and personate, tubular to salverform, tube base not spurred or gibbous, lobes 5, abaxial 3, adaxial 2, throat hairy;

stamens 2, medially adnate to corolla, filaments glabrous, pollen sacs parallel to filaments, connective not dilated;

staminodes 0 or 2, minute;

ovary 2-locular, placentation axile;

stigma crateriform.

Fruits

capsules, dehiscence septicidal, sometimes also loculicidal.

Capsules

conic, 4–5 × 2–2.8 mm.

Seeds

0.2–0.4 mm.

ca. 200, yellow to brown or black, ovoid to short-cylindric, wings absent.

2n

= 22.

Sophronanthe pilosa

Sophronanthe

Phenology Flowering May–Oct.
Habitat Sandy pinelands, bogs, swamps, cypress-gum depressions, pine savannas, wet flatwoods.
Elevation 0–700 m. (0–2300 ft.)
Distribution
from FNA
AL; AR; DE; FL; GA; KY; LA; MD; MS; NC; NJ; OK; SC; TN; TX; VA
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se United States
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Discussion

Plants with glabrous stems and leaves have been named Gratiola pilosa var. epilis. The combination for that taxon in Sophronanthe has not been made.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 2 (2 in the flora).

The two species of Sophronanthe often have been included in Gratiola or in the monospecific genera Sophronanthe (S. hispida) and Tragiola (S. pilosa). Molecular and morphological data support the monophyly of Sophronanthe (D. Estes 2008), which is sister to Gratiola in the narrow sense.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blades 0.7–3 mm wide, linear to lanceolate-ovate, margins prominently revolute; corollas salverform, 7–16 mm.
S. hispida
1. Leaf blades 4–11 mm wide, lanceolate to ovate, margins not or slightly revolute; corollas tubular, 5–9 mm.
S. pilosa
Source FNA vol. 17, p. 278. FNA vol. 17, p. 277. Author: Craig C. Freeman.
Parent taxa Plantaginaceae > Sophronanthe Plantaginaceae
Sibling taxa
S. hispida
Subordinate taxa
S. hispida, S. pilosa
Synonyms Gratiola pilosa, G. pilosa var. epilis, Tragiola pilosa Tragiola
Name authority (Michaux) Small: Fl. S.E. U.S., 1067, 1338. (1903) Bentham: in J. Lindley, Nat. Syst. Bot., 445. (1836)
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