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rough hedge-hyssop

hedge-hyssop

Habit Herbs, perennial; caudex thick, hard.
Stems

ascending to erect, 10–75 cm, villous.

ascending to erect, villous (hairs jointed), rarely glabrous.

Leaves

blade linear to lanceolate-ovate, 8–20 × 0.7–3 mm, base not clasping, truncate, margins entire, prominently revolute, apex acute to acuminate, surfaces villous, papillate, not distinctly punctate abaxially.

cauline, sometimes also basal, opposite;

petiole absent, sometimes present on basal leaves;

blade not fleshy, leathery, margins entire or serrate.

Inflorescences

axillary, flowers 1 or 2 per node;

bracts present.

Pedicels

0–1 mm;

bracteoles 6–9 mm.

present or absent;

bracteoles present, 2.

Flowers

calyx lobes 3–7 × 0.5–1 mm;

corolla salverform, 7–16 mm, tube white, veins inconspicuous or pale purple, limb white.

bisexual;

sepals 5, distinct, linear to linear-lanceolate, calyx ± bilaterally symmetric, campanulate;

corolla white, sometimes tinged purple or pink, bilaterally symmetric, bilabiate to bilabiate and personate, tubular to salverform, tube base not spurred or gibbous, lobes 5, abaxial 3, adaxial 2, throat hairy;

stamens 2, medially adnate to corolla, filaments glabrous, pollen sacs parallel to filaments, connective not dilated;

staminodes 0 or 2, minute;

ovary 2-locular, placentation axile;

stigma crateriform.

Fruits

capsules, dehiscence septicidal, sometimes also loculicidal.

Capsules

conic, 4–5 × 1.8–2.5 mm.

Seeds

0.2–0.4 mm.

ca. 200, yellow to brown or black, ovoid to short-cylindric, wings absent.

Sophronanthe hispida

Sophronanthe

Phenology Flowering May–Oct.
Habitat Sandy scrub, dry to wet flatwoods, sandhills, seasonally wet clearings.
Elevation 0–100 m. (0–300 ft.)
Distribution
from FNA
AL; FL; GA; MS
[BONAP county map]
se United States
[BONAP county map]
Discussion

Gratiola subulata Baldwin ex Bentham is an illegitimate name that pertains to Sophronanthe hispida.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 2 (2 in the flora).

The two species of Sophronanthe often have been included in Gratiola or in the monospecific genera Sophronanthe (S. hispida) and Tragiola (S. pilosa). Molecular and morphological data support the monophyly of Sophronanthe (D. Estes 2008), which is sister to Gratiola in the narrow sense.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blades 0.7–3 mm wide, linear to lanceolate-ovate, margins prominently revolute; corollas salverform, 7–16 mm.
S. hispida
1. Leaf blades 4–11 mm wide, lanceolate to ovate, margins not or slightly revolute; corollas tubular, 5–9 mm.
S. pilosa
Source FNA vol. 17, p. 278. FNA vol. 17, p. 277. Author: Craig C. Freeman.
Parent taxa Plantaginaceae > Sophronanthe Plantaginaceae
Sibling taxa
S. pilosa
Subordinate taxa
S. hispida, S. pilosa
Synonyms Gratiola hispida Tragiola
Name authority Bentham: in J. Lindley, Nat. Syst. Bot., 445. (1836) Bentham: in J. Lindley, Nat. Syst. Bot., 445. (1836)
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