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hedge-hyssop

shaggy hedge-hyssop

Habit Herbs, perennial; caudex thick, hard.
Stems

ascending to erect, villous (hairs jointed), rarely glabrous.

erect, 20–65 cm, villous, rarely glabrous.

Leaves

cauline, sometimes also basal, opposite;

petiole absent, sometimes present on basal leaves;

blade not fleshy, leathery, margins entire or serrate.

blade lanceolate to ovate, 8–20 × 4–13 mm, base clasping, rounded to subcordate, margins entire or serrate, not or slightly revolute, apex rounded to obtuse or acute, surfaces villous, rarely glabrous, papillate, distinctly punctate abaxially.

Inflorescences

axillary, flowers 1 or 2 per node;

bracts present.

Pedicels

present or absent;

bracteoles present, 2.

0–3(–5) mm;

bracteoles 6–9(–11) mm.

Flowers

bisexual;

sepals 5, distinct, linear to linear-lanceolate, calyx ± bilaterally symmetric, campanulate;

corolla white, sometimes tinged purple or pink, bilaterally symmetric, bilabiate to bilabiate and personate, tubular to salverform, tube base not spurred or gibbous, lobes 5, abaxial 3, adaxial 2, throat hairy;

stamens 2, medially adnate to corolla, filaments glabrous, pollen sacs parallel to filaments, connective not dilated;

staminodes 0 or 2, minute;

ovary 2-locular, placentation axile;

stigma crateriform.

calyx lobes 5–7 × 0.5–1.5 mm;

corolla tubular, 5–9 mm, tube yellow, veins bluish purple, limb white.

Fruits

capsules, dehiscence septicidal, sometimes also loculicidal.

Capsules

conic, 4–5 × 2–2.8 mm.

Seeds

ca. 200, yellow to brown or black, ovoid to short-cylindric, wings absent.

0.2–0.4 mm.

2n

= 22.

Sophronanthe

Sophronanthe pilosa

Phenology Flowering May–Oct.
Habitat Sandy pinelands, bogs, swamps, cypress-gum depressions, pine savannas, wet flatwoods.
Elevation 0–700 m. (0–2300 ft.)
Distribution
se United States
[BONAP county map]
from FNA
AL; AR; DE; FL; GA; KY; LA; MD; MS; NC; NJ; OK; SC; TN; TX; VA
[BONAP county map]
Discussion

Species 2 (2 in the flora).

The two species of Sophronanthe often have been included in Gratiola or in the monospecific genera Sophronanthe (S. hispida) and Tragiola (S. pilosa). Molecular and morphological data support the monophyly of Sophronanthe (D. Estes 2008), which is sister to Gratiola in the narrow sense.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Plants with glabrous stems and leaves have been named Gratiola pilosa var. epilis. The combination for that taxon in Sophronanthe has not been made.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blades 0.7–3 mm wide, linear to lanceolate-ovate, margins prominently revolute; corollas salverform, 7–16 mm.
S. hispida
1. Leaf blades 4–11 mm wide, lanceolate to ovate, margins not or slightly revolute; corollas tubular, 5–9 mm.
S. pilosa
Source FNA vol. 17, p. 277. Author: Craig C. Freeman. FNA vol. 17, p. 278.
Parent taxa Plantaginaceae Plantaginaceae > Sophronanthe
Sibling taxa
S. hispida
Subordinate taxa
S. hispida, S. pilosa
Synonyms Tragiola Gratiola pilosa, G. pilosa var. epilis, Tragiola pilosa
Name authority Bentham: in J. Lindley, Nat. Syst. Bot., 445. (1836) (Michaux) Small: Fl. S.E. U.S., 1067, 1338. (1903)
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