Sophronanthe |
Sophronanthe pilosa |
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hedge-hyssop |
shaggy hedge-hyssop |
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Habit | Herbs, perennial; caudex thick, hard. | |||||
Stems | ascending to erect, villous (hairs jointed), rarely glabrous. |
erect, 20–65 cm, villous, rarely glabrous. |
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Leaves | cauline, sometimes also basal, opposite; petiole absent, sometimes present on basal leaves; blade not fleshy, leathery, margins entire or serrate. |
blade lanceolate to ovate, 8–20 × 4–13 mm, base clasping, rounded to subcordate, margins entire or serrate, not or slightly revolute, apex rounded to obtuse or acute, surfaces villous, rarely glabrous, papillate, distinctly punctate abaxially. |
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Inflorescences | axillary, flowers 1 or 2 per node; bracts present. |
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Pedicels | present or absent; bracteoles present, 2. |
0–3(–5) mm; bracteoles 6–9(–11) mm. |
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Flowers | bisexual; sepals 5, distinct, linear to linear-lanceolate, calyx ± bilaterally symmetric, campanulate; corolla white, sometimes tinged purple or pink, bilaterally symmetric, bilabiate to bilabiate and personate, tubular to salverform, tube base not spurred or gibbous, lobes 5, abaxial 3, adaxial 2, throat hairy; stamens 2, medially adnate to corolla, filaments glabrous, pollen sacs parallel to filaments, connective not dilated; staminodes 0 or 2, minute; ovary 2-locular, placentation axile; stigma crateriform. |
calyx lobes 5–7 × 0.5–1.5 mm; corolla tubular, 5–9 mm, tube yellow, veins bluish purple, limb white. |
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Fruits | capsules, dehiscence septicidal, sometimes also loculicidal. |
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Capsules | conic, 4–5 × 2–2.8 mm. |
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Seeds | ca. 200, yellow to brown or black, ovoid to short-cylindric, wings absent. |
0.2–0.4 mm. |
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2n | = 22. |
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Sophronanthe |
Sophronanthe pilosa |
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Phenology | Flowering May–Oct. | |||||
Habitat | Sandy pinelands, bogs, swamps, cypress-gum depressions, pine savannas, wet flatwoods. | |||||
Elevation | 0–700 m. (0–2300 ft.) | |||||
Distribution | se United States |
AL; AR; DE; FL; GA; KY; LA; MD; MS; NC; NJ; OK; SC; TN; TX; VA |
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Discussion | Species 2 (2 in the flora). The two species of Sophronanthe often have been included in Gratiola or in the monospecific genera Sophronanthe (S. hispida) and Tragiola (S. pilosa). Molecular and morphological data support the monophyly of Sophronanthe (D. Estes 2008), which is sister to Gratiola in the narrow sense. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Plants with glabrous stems and leaves have been named Gratiola pilosa var. epilis. The combination for that taxon in Sophronanthe has not been made. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 277. | FNA vol. 17, p. 278. | ||||
Parent taxa | ||||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Tragiola | Gratiola pilosa, G. pilosa var. epilis, Tragiola pilosa | ||||
Name authority | Bentham: in J. Lindley, Nat. Syst. Bot., 445. (1836) | (Michaux) Small: Fl. S.E. U.S., 1067, 1338. (1903) | ||||
Web links |