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yellow necklacepod, yellow sophora

necklace-pod

Habit Shrubs or small trees, 1–3(–6) m, tomentulose to deeply sericeous. Herbs, perennial, shrubs, or trees, unarmed.
Stems

erect, pubescent or glabrous.

Leaves

rachis 10–25 cm;

leaflets 11–21, blades obovate, oblong, elliptic, or suborbiculate, 2–4(–5) cm, surfaces villous to sericeous abaxially, densely villous, sericeous, or glabrescent adaxially.

alternate, odd-pinnate;

stipules usually present, caducous, linear to deltate; petiolate, petiole 5–30 mm;

leaflets (7 or)9–23[–50], alternate or subopposite, stipels absent or minute and linear, blade margins entire, surfaces pubescent or glabrous (densely villous, sericeous, or glabrescent adaxially in S. tomentosa).

Inflorescences

35–75-flowered, crowded, 12–32 cm;

bracteoles 0.

5–75[+]-flowered, terminal or axillary, racemes or panicles;

bracts present, caducous;

bracteoles caducous or absent.

Pedicels

4–10 mm.

Flowers

spreading, (17–)20–25 mm;

calyx broadly campanulate, 5–8[–10] mm;

corolla creamy white to yellow;

ovary pubescent.

papilionaceous;

calyx campanulate, lobes 5, subequal, acute to truncate, sometimes gibbous, adaxial lobes often connate in part;

corolla white, creamy white, yellow, or purple, glabrous;

keel usually connate in part;

stamens 10, distinct or proximally connate;

anthers dorsifixed;

pistil linear to lanceolate.

Fruits

legumes, sessile or short-stipitate, narrowly oblong-moniliform, cylindric [compressed], fusiform, or torulose, dehiscent, pubescent [glabrous].

Legumes

black, narrowly oblong-moniliform, 5–15(–20) × 0.5–1 cm, leathery.

Seeds

1–15, light brown, 4–8 mm.

1–15, light brown to mustard-yellow, globose to subglobose.

x

= 9.

2n

= 18.

Sophora tomentosa

Sophora

Distribution
from USDA
sc United States; se United States; Mexico; Central America; West Indies; n South America; se Asia; Africa; Pacific Islands; Australia
[WildflowerSearch map]
[BONAP county map]
from USDA
United States; Mexico; Central America; South America; West Indies (Antilles, Bahamas); Eurasia; Africa; Pacific Islands; Australia
[BONAP county map]
Discussion

Subspecies ca. 7 (2 in the flora).

Sophora tomentosa is polymorphic and has a worldwide distribution along tropical and subtropical seashores. The seeds are considered toxic and have diuretic, sudorific, and purgative properties; they are also used for other medicinal purposes (R. A. Vines 1960).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species ca. 50 (4 in the flora).

Sophora as traditionally circumscribed is recognized as an unnatural assemblage. Molecular studies have elucidated the relationships among taxa in Sophora in the broad sense, resulting in recognition of several smaller genera. In the flora area, Sophora, as currently circumscribed, comprises herbaceous plants arising from a woody root, and the woody species S. tomentosa. Two species in the flora area that formerly were included in Sophora are now included in Dermatophyllum (D. arizonicum and D. secundiflorum), as is the closely related Mexican species D. gypsophilum and two others are placed in Styphnolobium (S. affine and S. japonicum). Styphnolobium is distinct from Sophora (in the narrow sense) in molecular studies, falling into the Cladrastis clade; Dermatophyllum is in a separate lineage from the Cladrastis clade and outside a clade in which the North American taxa of Sophora (in the narrow sense) fall (J. J. Doyle et al. 1996; R. T. Pennington et al. 2001; M. F. Wojciechowski et al. 2004).

The seeds and foliage of some species of Sophora contain neurotoxic alkaloids. Although neurotoxicity has not been demonstrated in native Sophora species, G. E. Burrows and R. L. Tyrl (2013) discussed possible teratogenic effects.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaflet blades mostly obovate or oblong; flowers (17–)20–23 mm.
subsp. bahamensis
1. Leaflet blades broadly elliptic or suborbiculate; flowers 23–25 mm.
subsp. occidentalis
1. Shrubs or small trees; corollas creamy white to yellow; coastal near beaches or in pinelands.
S. tomentosa
1. Herbs; corollas purple (fading blue), white, or creamy white; inland.
→ 2
2. Leaflet blades narrowly linear; leaf rachises 2–4 cm; corollas purple (fading blue).
S. stenophylla
2. Leaflet blades obovate to oblong or ovate to oblanceolate; leaf rachises (3–)5–17 cm; corollas white or creamy white.
→ 3
3. Inflorescences 7–15 cm; leaflet blades villous-tomentose abaxially, appressed- pubescent adaxially.
S. leachiana
3. Inflorescences 20–80 cm; leaflet blades sericeous abaxially, glabrous adaxially.
S. nuttalliana
Source FNA vol. 11. FNA vol. 11. Authors: Michael A. Vincent, Denis M. Kearns.
Parent taxa Fabaceae > subfam. Faboideae > Sophora Fabaceae > subfam. Faboideae
Sibling taxa
S. leachiana, S. nuttalliana, S. stenophylla
Subordinate taxa
S. tomentosa subsp. bahamensis, S. tomentosa subsp. occidentalis
S. leachiana, S. nuttalliana, S. stenophylla, S. tomentosa
Synonyms Pseudosophora, Radiusia, Vexibia, Zanthyrsis
Name authority Linnaeus: Sp. Pl. 1: 373. (1753) Linnaeus: Sp. Pl. 1: 373. (1753): Gen. Pl. ed. 5, 175. (1754)
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