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California goldenrod, three-nerve goldenrod, velvety goldenrod

common wrinkle-leaf goldenrod, rough-stem goldenrod, verge d'or rugueuse, wrinkle-leaf goldenrod

Habit Plants (forming diffuse clones) 15–80 (–150) cm; rhizomes creeping, slender. Plants 30–200 cm; rhizomes long-creeping, forming clones.
Stems

1 (at ends of rhizomes), ascending to erect, glabrate proximally to sparsely to densely strigoso-puberulent distally.

1–50+, erect, glabrous or densely hispid to strigose.

Leaves

basal and proximal cauline often persisting to flowering, gradually tapering to winged petioles, blades linear oblanceolate to oblanceolate, rarely spatulate, 50–120 × 8–30 mm, proximalmost much smaller, margins entire to sharply serrate, faces glabrate to moderately scabroso-strigose;

mid and distal cauline sessile or subsessille, blades elliptic to oblanceolate or obovate, 10–50 × 3–12 mm, mid tapering to bases, somewhat to strongly 3-nerved, largest, usually much reduced distally, margins entire or sometimes distally serrate, apices acute, faces sparsely to densely strigoso-puberulent, sometimes softly so.

basal withering by flowering;

proximal cauline usually withering by flowering, sessile, blades elliptic to lanceolate, 68–104 × 20–25 mm, margins sharply serrate, apices acute to attenuate, abaxial faces usually hispido-strigose (at least on main nerves), nerves sometimes prominent, abaxial glabrate;

mid to distal cauline sessile, blades lanceolate, elliptic or ovate, (15–)40–75(–90) × (6–)12–22(–32) mm, largest at midstem, somewhat reduced to much reduced distally, margins coarsely to finely serrate, ciliate, indument similar to proximal or denser.

Peduncles

1–6 mm, sparsely to densely strigillose;

bracteoles 0–5, sometimes clustered near to and grading into phyllaries, linear-lanceolate.

1–1.8 mm, sparsely to densely hispido-strigillose;

bracteoles linear- lanceolate to ovate.

Involucres

campanulate, 3.5–6 mm.

narrowly campanulate, (2–)2.5–3.5(–4.5) mm.

Ray florets

6–12;

laminae 2.9–6.3 × 0.3–0.7(–1) mm.

(4–)6–8(–12);

laminae (0.9–)1–1.6(–2.3) × 0.4–0.7 mm.

Disc florets

5–17;

corollas 3.5–6 mm, lobes 0.8–1.7 mm.

(2–)4–6(–8);

corollas 2–3.5(–4.5) mm, lobes (0.5–)0.7–1(–1.3) mm.

Phyllaries

in 3–5 series, lanceolate to oblong, strongly unequal, acute or sometimes obtuse, glabrous or sparsely strigillose.

in 3–4 series, strongly unequal, lanceolate to linear-lanceolate, acute to obtuse.

Heads

(2–)30–500, in narrow to broad, thyrsiform to secund-pyramidal paniculiform arrays, branches recurved, secund, congested to lax.

50–1500, secund, in secund-pyramidal paniculiform arrays 7–36(–50) × 9–26 cm, compact to lax, branches divergent and recurved, longest 0.8–34 cm, leafy-bracteate.

Cypselae

0.7–2.7 mm, sparsely to densely strigillose;

pappi 2.5–4.7 mm.

(narrowly obconic) 0.9–1.5 mm, moderately strigillose;

pappi 1.8–2.5 mm.

Solidago velutina

Solidago rugosa

Distribution
from FNA
AZ; CA; CO; ID; NM; NV; OR; TX; WY; Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DE; FL; GA; IL; IN; KY; LA; MA; MD; ME; MI; MO; MS; NC; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; VT; WI; WV; NB; NS; ON; PE; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 3 (2 in the flora).

G. L. Nesom (1993b) merged Solidago californica, S. sparsiflora, and S. velutina without recognizing any infraspecific taxa, as did A. Cronquist (1994). J. C. Semple et al. (1990) compared S. californica and S. sparsiflora to S. nemoralis and found that all three are significantly different in a multivariate analysis. Evidence for separating the two subspecies of S. nemoralis was greater than the support for separating S. californica and S. sparsiflora.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

Solidago rugosa is highly variable in size, array shape, and hairiness. It is similar to members of the S. canadensis complex; it differs in not having 3-nerved leaves. The species is divided into two subspecies and five varieties that can be difficult to distinguish.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems and leaves sparsely strigillose; outer phyllaries glabrate; Wyoming to e California, s to Arizona and New Mexico, Mexico
subsp. sparsiflora
1. Stems and leaves moderately to densely strigillose; outer phyllaries sparsely strigillose; California and s Oregon
subsp. californica
1. Leaves relatively thin, not very rugose, usually sharply toothed, apices acuminate, glabrous or relatively soft-hairy; ray florets (4–)6–11(–13); northern (subsp. rugosa)
→ 2
1. Leaves relatively thick and firm, strongly rugose-nerved, usually blunt-toothed to subentire, apices often acute, relatively short and stiff hairy; ray florets 4–9; mostly se United States (subsp. aspera)
→ 3
2. Stems and leaves hairy; e Canada and ne United States s to Virginia
var. rugosa
2. Stems and leaves glabrous; coastal cedar bogs and swamps
var. sphagnophila
3. Arrays narrow, proximal branches not much exceeding subtending leaves; herbage sparsely hairy; mid to higher elevations in mountains
var. cronquistiana
3. Arrays wide, usually with elongate proximal branches greatly exceeding subtending leaves; herbage moderately to densely hairy; lower elevations in mountains, piedmont, and coastal plain
→ 4
4. Distal cauline leaves lanceolate to elliptic, not much reduced distally; much of range of subspecies
var. aspera
4. Distal cauline leaves ovate, much reduced distally; outer coastal plain
var. celtidifolia
Source FNA vol. 20, p. 160. FNA vol. 20, p. 146.
Parent taxa Asteraceae > tribe Astereae > Solidago > sect. Solidago > subsect. Nemorales Asteraceae > tribe Astereae > Solidago > sect. Solidago > subsect. Venosae > ser. Venosae
Sibling taxa
S. albopilosa, S. altiplanities, S. altissima, S. arenicola, S. arguta, S. auriculata, S. bicolor, S. brachyphylla, S. buckleyi, S. caesia, S. canadensis, S. confinis, S. curtisii, S. delicatula, S. drummondii, S. elongata, S. erecta, S. faucibus, S. fistulosa, S. flexicaulis, S. gattingeri, S. gigantea, S. glomerata, S. guiradonis, S. hispida, S. houghtonii, S. juliae, S. juncea, S. kralii, S. lancifolia, S. latissimifolia, S. leavenworthii, S. leiocarpa, S. lepida, S. ludoviciana, S. macrophylla, S. missouriensis, S. mollis, S. multiradiata, S. nana, S. nemoralis, S. nitida, S. odora, S. ohioensis, S. ouachitensis, S. patula, S. petiolaris, S. pinetorum, S. plumosa, S. ptarmicoides, S. puberula, S. pulchra, S. radula, S. riddellii, S. rigida, S. roanensis, S. rugosa, S. rupestris, S. sciaphila, S. sempervirens, S. shortii, S. simplex, S. spathulata, S. speciosa, S. spectabilis, S. sphacelata, S. spithamaea, S. squarrosa, S. stricta, S. tarda, S. tortifolia, S. uliginosa, S. ulmifolia, S. verna, S. villosicarpa, S. wrightii
S. albopilosa, S. altiplanities, S. altissima, S. arenicola, S. arguta, S. auriculata, S. bicolor, S. brachyphylla, S. buckleyi, S. caesia, S. canadensis, S. confinis, S. curtisii, S. delicatula, S. drummondii, S. elongata, S. erecta, S. faucibus, S. fistulosa, S. flexicaulis, S. gattingeri, S. gigantea, S. glomerata, S. guiradonis, S. hispida, S. houghtonii, S. juliae, S. juncea, S. kralii, S. lancifolia, S. latissimifolia, S. leavenworthii, S. leiocarpa, S. lepida, S. ludoviciana, S. macrophylla, S. missouriensis, S. mollis, S. multiradiata, S. nana, S. nemoralis, S. nitida, S. odora, S. ohioensis, S. ouachitensis, S. patula, S. petiolaris, S. pinetorum, S. plumosa, S. ptarmicoides, S. puberula, S. pulchra, S. radula, S. riddellii, S. rigida, S. roanensis, S. rupestris, S. sciaphila, S. sempervirens, S. shortii, S. simplex, S. spathulata, S. speciosa, S. spectabilis, S. sphacelata, S. spithamaea, S. squarrosa, S. stricta, S. tarda, S. tortifolia, S. uliginosa, S. ulmifolia, S. velutina, S. verna, S. villosicarpa, S. wrightii
Subordinate taxa
S. velutina subsp. californica, S. velutina subsp. sparsiflora
S. rugosa var. aspera, S. rugosa var. celtidifolia, S. rugosa var. cronquistiana, S. rugosa var. rugosa, S. rugosa var. sphagnophila
Synonyms Aster velutinus Aster rugosus
Name authority de Candolle: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 5: 332. (1836) Miller: Gard. Dict. ed. 8, Solidago no. 25. (1768)
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