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twist-leaf goldenrod

common wrinkle-leaf goldenrod, rough-stem goldenrod, verge d'or rugueuse, wrinkle-leaf goldenrod

Habit Plants 30–130 cm; caudices small, woody, rhizomes creeping, elongate. Plants 30–200 cm; rhizomes long-creeping, forming clones.
Stems

1–10, ascending to erect, uniformly finely strigillose-villous distal to mid.

1–50+, erect, glabrous or densely hispid to strigose.

Leaves

basal cauline nearly always withering well before flowering, linear-oblanceolate, smaller than proximal mid cauline, serrate;

proximal persisting, sometimes brown-black after senescence, often twisted;

proximal to distal numerous (100+ on tall stems), crowded, sessile, blades (bright green) linear to linear-lanceolate, 20–70 × 2–7(–10) mm, margins remotely serrulate (proximal) to entire or with 1–2 minute serrations (distal), obscurely 3-nerved, one or both faces glabrous or finely strigillose.

basal withering by flowering;

proximal cauline usually withering by flowering, sessile, blades elliptic to lanceolate, 68–104 × 20–25 mm, margins sharply serrate, apices acute to attenuate, abaxial faces usually hispido-strigose (at least on main nerves), nerves sometimes prominent, abaxial glabrate;

mid to distal cauline sessile, blades lanceolate, elliptic or ovate, (15–)40–75(–90) × (6–)12–22(–32) mm, largest at midstem, somewhat reduced to much reduced distally, margins coarsely to finely serrate, ciliate, indument similar to proximal or denser.

Peduncles

1–4 mm, finely strigilloso-villous;

bracteoles 0–2, linear, grading into phyllaries.

1–1.8 mm, sparsely to densely hispido-strigillose;

bracteoles linear- lanceolate to ovate.

Involucres

narrowly campanulate, 2.5–3.5 mm.

narrowly campanulate, (2–)2.5–3.5(–4.5) mm.

Ray florets

2–8;

laminae 1–2 × 0.25–0.5 mm.

(4–)6–8(–12);

laminae (0.9–)1–1.6(–2.3) × 0.4–0.7 mm.

Disc florets

2–4(–6);

corollas 2.3–3.5 mm, lobes 0.4–1 mm.

(2–)4–6(–8);

corollas 2–3.5(–4.5) mm, lobes (0.5–)0.7–1(–1.3) mm.

Phyllaries

in 3–4 series, strongly unequal;

outer ovate, acute, inner oblong, obtuse to rounded.

in 3–4 series, strongly unequal, lanceolate to linear-lanceolate, acute to obtuse.

Heads

100–300+, in short to elongate, pyramidal paniculiform arrays, branches recurved, secund.

50–1500, secund, in secund-pyramidal paniculiform arrays 7–36(–50) × 9–26 cm, compact to lax, branches divergent and recurved, longest 0.8–34 cm, leafy-bracteate.

Cypselae

(narrowly obconic) 1 mm, strigillose;

pappi 2–3 mm.

(narrowly obconic) 0.9–1.5 mm, moderately strigillose;

pappi 1.8–2.5 mm.

2n

= 18.

Solidago tortifolia

Solidago rugosa

Phenology Flowering Aug–Nov (year-round).
Habitat Dry, usually sandy soils, pinelands, embankments
Elevation 0–100+ m (0–300+ ft)
Distribution
from FNA
AL; AR; FL; GA; LA; MS; NC; SC; TX; VA
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DE; FL; GA; IL; IN; KY; LA; MA; MD; ME; MI; MO; MS; NC; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; VT; WI; WV; NB; NS; ON; PE; QC
[WildflowerSearch map]
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

Solidago rugosa is highly variable in size, array shape, and hairiness. It is similar to members of the S. canadensis complex; it differs in not having 3-nerved leaves. The species is divided into two subspecies and five varieties that can be difficult to distinguish.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves relatively thin, not very rugose, usually sharply toothed, apices acuminate, glabrous or relatively soft-hairy; ray florets (4–)6–11(–13); northern (subsp. rugosa)
→ 2
1. Leaves relatively thick and firm, strongly rugose-nerved, usually blunt-toothed to subentire, apices often acute, relatively short and stiff hairy; ray florets 4–9; mostly se United States (subsp. aspera)
→ 3
2. Stems and leaves hairy; e Canada and ne United States s to Virginia
var. rugosa
2. Stems and leaves glabrous; coastal cedar bogs and swamps
var. sphagnophila
3. Arrays narrow, proximal branches not much exceeding subtending leaves; herbage sparsely hairy; mid to higher elevations in mountains
var. cronquistiana
3. Arrays wide, usually with elongate proximal branches greatly exceeding subtending leaves; herbage moderately to densely hairy; lower elevations in mountains, piedmont, and coastal plain
→ 4
4. Distal cauline leaves lanceolate to elliptic, not much reduced distally; much of range of subspecies
var. aspera
4. Distal cauline leaves ovate, much reduced distally; outer coastal plain
var. celtidifolia
Source FNA vol. 20, p. 158. FNA vol. 20, p. 146.
Parent taxa Asteraceae > tribe Astereae > Solidago > sect. Solidago > subsect. Triplinerviae Asteraceae > tribe Astereae > Solidago > sect. Solidago > subsect. Venosae > ser. Venosae
Sibling taxa
S. albopilosa, S. altiplanities, S. altissima, S. arenicola, S. arguta, S. auriculata, S. bicolor, S. brachyphylla, S. buckleyi, S. caesia, S. canadensis, S. confinis, S. curtisii, S. delicatula, S. drummondii, S. elongata, S. erecta, S. faucibus, S. fistulosa, S. flexicaulis, S. gattingeri, S. gigantea, S. glomerata, S. guiradonis, S. hispida, S. houghtonii, S. juliae, S. juncea, S. kralii, S. lancifolia, S. latissimifolia, S. leavenworthii, S. leiocarpa, S. lepida, S. ludoviciana, S. macrophylla, S. missouriensis, S. mollis, S. multiradiata, S. nana, S. nemoralis, S. nitida, S. odora, S. ohioensis, S. ouachitensis, S. patula, S. petiolaris, S. pinetorum, S. plumosa, S. ptarmicoides, S. puberula, S. pulchra, S. radula, S. riddellii, S. rigida, S. roanensis, S. rugosa, S. rupestris, S. sciaphila, S. sempervirens, S. shortii, S. simplex, S. spathulata, S. speciosa, S. spectabilis, S. sphacelata, S. spithamaea, S. squarrosa, S. stricta, S. tarda, S. uliginosa, S. ulmifolia, S. velutina, S. verna, S. villosicarpa, S. wrightii
S. albopilosa, S. altiplanities, S. altissima, S. arenicola, S. arguta, S. auriculata, S. bicolor, S. brachyphylla, S. buckleyi, S. caesia, S. canadensis, S. confinis, S. curtisii, S. delicatula, S. drummondii, S. elongata, S. erecta, S. faucibus, S. fistulosa, S. flexicaulis, S. gattingeri, S. gigantea, S. glomerata, S. guiradonis, S. hispida, S. houghtonii, S. juliae, S. juncea, S. kralii, S. lancifolia, S. latissimifolia, S. leavenworthii, S. leiocarpa, S. lepida, S. ludoviciana, S. macrophylla, S. missouriensis, S. mollis, S. multiradiata, S. nana, S. nemoralis, S. nitida, S. odora, S. ohioensis, S. ouachitensis, S. patula, S. petiolaris, S. pinetorum, S. plumosa, S. ptarmicoides, S. puberula, S. pulchra, S. radula, S. riddellii, S. rigida, S. roanensis, S. rupestris, S. sciaphila, S. sempervirens, S. shortii, S. simplex, S. spathulata, S. speciosa, S. spectabilis, S. sphacelata, S. spithamaea, S. squarrosa, S. stricta, S. tarda, S. tortifolia, S. uliginosa, S. ulmifolia, S. velutina, S. verna, S. villosicarpa, S. wrightii
Subordinate taxa
S. rugosa var. aspera, S. rugosa var. celtidifolia, S. rugosa var. cronquistiana, S. rugosa var. rugosa, S. rugosa var. sphagnophila
Synonyms Aster rugosus
Name authority Elliott: Sketch Bot. S. Carolina 2: 377. (1823) Miller: Gard. Dict. ed. 8, Solidago no. 25. (1768)
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