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noble goldenrod, showy goldenrod

goldenrod

Habit Plants (30–)50–200 cm; caudices stout, woody. Perennials, 5–100(–200) cm; woody caudices or rhizomes.
Stems

1(–5), erect, glabrous proximally to strigillose in arrays.

decumbent to ascending or erect, sometimes branched distally, glabrous or strigose, strigillose, hispid, or short-villous.

Leaves

basal and proximal cauline (sometimes withering by flowering) tapering to winged petioles, blades lanceolate to ovate-elliptic, 50–300 (including petiole) × 12–80 mm (sometimes firm), margins sharply serrate to crenate or entire, faces glabrate to sparsely strigillose;

mid to distal cauline sessile, blades narrowly ovate to lanceolate or elliptic, 25–90 × 5–30 mm, gradually reduced distally, margins serrulate to entire (distally), ciliate, faces glabrous or sparsely scabroso-strigose.

basal (persistent or not by flowering) and cauline; petiolate (proximal) or sessile (proximal and distal, latter sometimes subpetiolate);

proximal blades sometimes 3-nerved, ovate-oblanceolate, margins often serrate, faces glabrous or densely hairy;

distal sometimes 3-nerved, glabrous or sparsely to densely scabrous, strigillose, or villous, sometimes stipitate-glandular, sometimes resinous.

Peduncles

1.5–3 mm, sparsely to moderately scabroso-strigillose;

bracteoles linear, grading into phyllaries, scattered along peduncles, clustered near heads.

Involucres

narrowly campanulate, 4–6.5 mm.

campanulate to cylindric (often spreading upon drying), 3–12 × 1.7–10 mm.

Receptacles

slightly convex, pitted, epaleate.

Ray florets

(2–)3–7(–9);

laminae 3–4 × 0.5–1 mm.

(0–)2–15(–24), pistillate, fertile;

corollas yellow, rarely white (usually glabrous).

Disc florets

6–16;

corollas 2.5–4 mm, lobes 0.5–1.2 mm.

2–35(–60), bisexual, fertile;

corollas yellow, ± ampliate, tubes shorter than throats (usually glabrous), lobes 5, erect to spreading, triangular to narrowly lanceolate;

style-branch appendages broadly to narrowly triangular (lengths 0.7–1 times stigmatic lines).

Phyllaries

in 3–4 series, appressed, strongly unequal, outer ovate, mid and inner lanceolate, (midnerves often raised and thick) apices acute to obtuse or rounded, glabrous.

10–35 in 3–5 series, midnerves usually ± swollen and translucent, sometimes plus 2–5 secondary nerves (striate, flat), linear-lanceolate to oblong or ovate, unequal to rarely subequal, margins scarious, (apices rounded to acute or attenuate), faces glabrous or sparsely pilose or puberulent, sometimes minutely stipitate-glandular, sometimes resinous.

Heads

15–300+, not secund, in usually dense, sometimes open, elongate, paniculiform to thyrsiform arrays, (5–)10–45 × (2–)3–7(–12) cm;

branches strongly ascending, often racemiform.

usually radiate, sometimes discoid, (1–)2–1500+ in racemiform (club-shaped or pyramidal), paniculiform or corymbo-paniculiform, sometimes secund arrays.

Cypselae

(narrowly obconic) 1.6–2.5 mm, glabrous;

pappi 3–4.5 mm.

narrowly obconic to cylindric, sometimes somewhat compressed, ribs usually 8–10 (either darker and sometimes translucent or lighter than bodies), glabrous or moderately strigillose;

pappi persistent, outer setiform scales (0.25–0.5 mm; rarely present) plus 2 series of 25–45 longer, barbellate bristles, mid apically attenuate, 90–95% length of inner, inner apically weakly to strongly clavate [(0.5–in S. sphacelata)1.5–5(–7.3) mm].

x

= 9.

Solidago speciosa

Solidago

Distribution
from FNA
AL; AR; CO; CT; DE; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MN; MO; MS; NC; ND; NE; NJ; NM; NY; OH; OK; RI; SC; SD; TN; TX; VA; VT; WI; WV; WY; ON
[WildflowerSearch map]
[BONAP county map]
from USDA
North America; Mexico; South America; Eurasia
[BONAP county map]
Discussion

Subspecies 2 (2 in the flora).

Solidago speciosa occurs scattered throughout its range, often in small populations. Five races have been acknowledged; only three varieties appear to warrant recognition. The species is divided into two nearly allopatric subspecies. The typical subspecies includes two varieties that are sometimes difficult to distinguish and considerably overlap in their ranges. The differences in size and number of leaves and persistence of the proximal cauline leaves may be caused in part by growing conditions.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species ca. 100 (77 in the flora).

Solidago is found primarily in North America with some South American and Eurasian species (8 in Mexico, 4 in South America, 6–10 in Europe and Asia).

Recent studies on the phylogeny of Astereae and Solidago have done much to resolve relationships of the goldenrods to each other and to other genera within the tribe (J. J. Zhang 1996; J. C. Semple et al. 1999; R. D. Noyes and L. H. Rieseberg 1999; J. B. Beck et al. 2004). Traditionally the grass-leaved goldenrods, Euthamia, have been included in Solidago; they are well separated within the North American clade. L. C. Anderson and J. B. Creech (1975) presented anatomic reasons for separating the two genera and also Chrysoma, Gundlachia, and Petradoria. The segregate genera Oligoneuron and Brachychaeta fit within Solidago based on anatomy and DNA studies. Brintonia lies outside Solidago based on DNA and morphology. The position of the monophyletic Oreochrysum also appears to lie outside and phylogenetically close to Solidago. G. L. Nesom (1993) placed the flat-topped goldenrods in Oligoneuron. Results from Zhang’s study indicate that the flat-topped goldenrods are nested within other goldenrod groups that Nesom included in Solidago. Beck et al. (2004) indicated that further work is needed before a definitive molecular-based phylogeny is available. While the flat-topped goldenrods are readily distinct from other goldenrods on the basis of arrangements of heads, additional features such as double pappus with clavate inner bristles and oligoneurate phyllaries occur in other species of Solidago as well. The flat-topped goldenrods are treated here as Solidago sect. Ptarmicoidei following Semple and K. N. Gandhi (2004). Nesom provided much of the formal subgeneric nomenclature that by and large is followed here; there are some differences in assignment of species to subsections. Semple (2003, 2004) proposed a number of additional new names and combinations used here. J. L. A. Hood and Semple (2003) presented evidence that the pappus of Solidago is not simple, as previously assumed, but is like that of related genera: it is in two series, inner series somewhat clavate and slightly longer, or sometimes in three series with an additional, shorter outer series of few setiform scales.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves pale green, somewhat glaucous; plants 30–80 cm; e Colorado, extreme w Oklahoma, and adjacent New Mexico, disjunct in Black Hills (S. speciosa subsp. pallida)
var. pallida
1. Leaves usually dark green, not glaucous; plants usually 80–200 cm; e deciduous forest onto e prairies and oak savannas (S. speciosa subsp. speciosa)
→ 2
2. Basal leaves usually present at flowering, (20–) 28–55 mm wide, often coarsely serrate; mid cauline leaves (10–)20–28 mm wide, not crowded, not stiff, not scabrous; e UnitedStates
var. speciosa
2. Basal leaves persisting or not to flowering, 8–20 mm wide, often entire or shallowly serrate; mid cauline leaves 4–15(–20) mm wide, often crowded, usually stiff, often somewhat scabrous; w portions of deciduous forest and adjacent prairies, glade habitats further east
var. rigidiuscula
1. Heads usually in ± corymbiform (flat-topped or rounded) arrays, sometimes in glomerules
→ 2
1. Heads in ± paniculiform or thyrsiform arrays (wand-, club-, or secund cone-shaped) or in axillary clusters
→ 3
2. Bases of basal leaves not persistent to flowering; heads in round-topped arrays; phyllaries not striate; arctic and alpine habitats, Canada, Alaska, s to California and New Mexico in Western Cordillera, exposed rock outcrops at 1500–2000 m in s Appala-chians
(sect. Solidago) subsect. Multiradiatae
2. Bases of basal leaves persistent; heads in flat-topped arrays (somewhat rounded in S. riddellii), sometimes also in glomerules; phyllaries usually striate (except in S. ptarmicoides); Great Plains, Great Lakes region, glades and savannas in e United States
sect. Ptarmicoidei
3. Pappi less than 1/4 length of disc corollas; c, se United States
(sect. Solidago) ser. Brachychaeta
3. Pappi more than 1/2 length of disc corollas; much of North America
→ 4
4. Cauline leaves: bases of petioles or blades usually cordate and auriculate-clasping; woods, se United States
(sect. Solidago) ser. Auriculatae
4. Cauline leaves: bases of petioles or blades not cordate and auriculate-clasping; various habitats across continent
→ 5
5. Heads in axillary clusters or in thyrsiform arrays, not nodding or secund at summit nor with recurved, secund branches
→ 6
5. Heads in paniculiform arrays, nodding or secund distally and/or with proximal branches ± recurved and/or secund
→ 10
6. Basal and proximalmost cauline leaves usually withering by flowering or relatively small
→ 7
6. Basal and proximalmost cauline leaves present at flowering and relatively large
→ 8
7. Phyllaries sparsely to densely stipitate-glandular; leaves sometimes resinous or stipitate-glandular; mid to distal cauline leaves usually petiolate (petioles ca. 1–2 mm)
(sect. Solidago) subsect. Thyrsiflorae
7. Phyllaries not stipitate-glandular; leaves not stipitate-glandular or resin-ous; distal cauline leaves sessile
(sect. Solidago) subsect. Glomeruliflorae
8. Phyllaries, peduncles, and/or leaves ± viscid (glands minute); mid to dis- tal cauline leaves petiolate
(sect. Solidago) subsect. Humiles
8. Phyllaries, peduncles, and leaves usually not viscid (glands on phyllaries in S. villosicarpa); mid to distal cauline leaves sessile
→ 9
9. Petioles of proximal cauline leaves not sheathing stems, leaves not thick and somewhat fleshy; usually mesic habitats
(sect. Solidago) subsect. Squarrosae
9. Petioles of proximal cauline leaves sheathing stems, leaves often thick and somewhat fleshy; bog and marsh habitats
(sect. Solidago) subsect. Maritimae
10. Basal and proximalmost cauline leaves present at flowering and relatively large; mid to distal cauline leaves much reduced distally.
→ 11
10. Basal and proximalmost cauline leaves usually withering by flowering or relatively small; mid to distal cauline leaves gradually reduced distally, similar to proximal
→ 14
11. Blades of basal and proximal cauline leaves ovate to lanceolate
(sect. Solidago) ser. Argutae
11. Blades of basal and proximal cauline leaves lanceolate to linear-lanceolate (basal blades sometimes ovate in S. nemoralis)
→ 12
12. Petiole bases of proximalmost leaves sheathing stems; plantsof marshes and bogs
(sect. Solidago) subsect. Maritimae
12. Petiole bases of proximalmost leaves not sheathing stems; usually mesic to drier habitats
→ 13
13. Stems and leaves sparsely to densely hairy
(sect. Solidago) subsect. Nemorales
13. Stems and leaves usually glabrous
Solidago (sect. Solidago) subsect. Junceae
14. Plants usually glabrous (even among heads)
Solidago (sect. Solidago) subsect. Junceae
14. Plants puberulent (at least among heads).
→ 15
15. Blades of mid cauline leaves reticulate-nerved with multiple prominent lateral nerves abaxially (lateral nerves usually prominent only on proximal cauline leaves in S. fistulosa and S. odora), not 3-nerved, obovate, elliptic, lanceolate, oblanceolate but usually not linear (if linear, anise-scented when crushed)
(sect. Solidago) subsect. Venosae
15. Blades of mid cauline leaves usually 3-nerved (lateral nerves sometimes obscured in S. altiplanities, S. leavenworthii, and S. tortifolia, and in some plants of S. lepida from n Rocky Mountains), narrowly to broadly ovate, elliptic, lanceolate, oblanceolate, or linear
→ 16
16. Blades of mid cauline leaves narrowly to broadly ovate; woods, e Missouri, adjacent Arkansas, Illinois, Kentucky
(sect. Solidago) ser. Drummondiani
16. Blades of mid cauline leaves elliptic, lanceolate, oblanceolate, or linear; various habitats across the continent (see also 163a.11. Subsect. Nemorales for plants with only distalcauline leaves present)
(sect. Solidago) subsect. Triplinerviae
Source FNA vol. 20, p. 123. FNA vol. 20, p. 107. Authors: John C. Semple, Rachel E. Cook.
Parent taxa Asteraceae > tribe Astereae > Solidago > sect. Solidago > subsect. Squarrosae Asteraceae > tribe Astereae
Sibling taxa
S. albopilosa, S. altiplanities, S. altissima, S. arenicola, S. arguta, S. auriculata, S. bicolor, S. brachyphylla, S. buckleyi, S. caesia, S. canadensis, S. confinis, S. curtisii, S. delicatula, S. drummondii, S. elongata, S. erecta, S. faucibus, S. fistulosa, S. flexicaulis, S. gattingeri, S. gigantea, S. glomerata, S. guiradonis, S. hispida, S. houghtonii, S. juliae, S. juncea, S. kralii, S. lancifolia, S. latissimifolia, S. leavenworthii, S. leiocarpa, S. lepida, S. ludoviciana, S. macrophylla, S. missouriensis, S. mollis, S. multiradiata, S. nana, S. nemoralis, S. nitida, S. odora, S. ohioensis, S. ouachitensis, S. patula, S. petiolaris, S. pinetorum, S. plumosa, S. ptarmicoides, S. puberula, S. pulchra, S. radula, S. riddellii, S. rigida, S. roanensis, S. rugosa, S. rupestris, S. sciaphila, S. sempervirens, S. shortii, S. simplex, S. spathulata, S. spectabilis, S. sphacelata, S. spithamaea, S. squarrosa, S. stricta, S. tarda, S. tortifolia, S. uliginosa, S. ulmifolia, S. velutina, S. verna, S. villosicarpa, S. wrightii
Subordinate taxa
S. speciosa var. pallida, S. speciosa var. rigidiuscula, S. speciosa var. speciosa
Solidago (sect. Solidago) ser. Argutae, Solidago (sect. Solidago) ser. Auriculatae, Solidago (sect. Solidago) ser. Brachychaeta, Solidago (sect. Solidago) ser. Drummondiani, Solidago (sect. Solidago) subsect. Glomeruliflorae, Solidago (sect. Solidago) subsect. Humiles, Solidago (sect. Solidago) subsect. Maritimae, Solidago (sect. Solidago) subsect. Multiradiatae, Solidago (sect. Solidago) subsect. Nemorales, Solidago (sect. Solidago) subsect. Squarrosae, Solidago (sect. Solidago) subsect. Thyrsiflorae, Solidago (sect. Solidago) subsect. Triplinerviae, Solidago (sect. Solidago) subsect. Venosae, S. sect. Ptarmicoidei
Synonyms Aster speciosus Actipsis, Aster subg. S., Leioligo, Oligoneuron
Name authority Nuttall: Gen. N. Amer. Pl. 2: 160. (1818) Linnaeus: Sp. Pl. 2: 878. (1753): Gen. Pl. ed. 5, 374. (1754)
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