Solidago simplex |
Solidago velutina |
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alpine goldenrod, dune goldenrod, Mt. Albert goldenrod, Rand's goldenrod, spikelike goldenrod, sticky goldenrod |
California goldenrod, three-nerve goldenrod, velvety goldenrod |
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Habit | Plants 5–80 cm; caudices branching. | Plants (forming diffuse clones) 15–80 (–150) cm; rhizomes creeping, slender. | ||||||||||||||||||||||||||||||||
Stems | 1–10+, ascending to erect, proximally glabrous, strigose in arrays. |
1 (at ends of rhizomes), ascending to erect, glabrate proximally to sparsely to densely strigoso-puberulent distally. |
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Leaves | basal and proximal cauline petiolate, blades narrowly oblanceolate, (20–)50–100(–160) × (2–)5–16(–56) mm, margins serrate to crenate, apices acute to obtuse, glabrous; mid and distal sessile, similar, blades lanceolate to linear, 12–45 × 2–19 mm, reduced distally, margins entire to sparsely serrate, sometimes resinous. |
basal and proximal cauline often persisting to flowering, gradually tapering to winged petioles, blades linear oblanceolate to oblanceolate, rarely spatulate, 50–120 × 8–30 mm, proximalmost much smaller, margins entire to sharply serrate, faces glabrate to moderately scabroso-strigose; mid and distal cauline sessile or subsessille, blades elliptic to oblanceolate or obovate, 10–50 × 3–12 mm, mid tapering to bases, somewhat to strongly 3-nerved, largest, usually much reduced distally, margins entire or sometimes distally serrate, apices acute, faces sparsely to densely strigoso-puberulent, sometimes softly so. |
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Peduncles | 3.1–10.3 mm, glabrate to sparsely strigillose; bracteoles few, linear. |
1–6 mm, sparsely to densely strigillose; bracteoles 0–5, sometimes clustered near to and grading into phyllaries, linear-lanceolate. |
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Involucres | campanulate, 3–7 mm. |
campanulate, 3.5–6 mm. |
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Ray florets | 7–16; laminae 2–5 × 0.7–0.9 mm. |
6–12; laminae 2.9–6.3 × 0.3–0.7(–1) mm. |
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Disc florets | 6–31; corollas 4–4.9 mm, lobes 0.6–1.3(–2) mm. |
5–17; corollas 3.5–6 mm, lobes 0.8–1.7 mm. |
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Phyllaries | (in 3–4 series) strongly unequal, often resinous; outer ovate, acute, inner linear-oblong, obtuse. |
in 3–5 series, lanceolate to oblong, strongly unequal, acute or sometimes obtuse, glabrous or sparsely strigillose. |
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Heads | 3–150, not secund, in narrowly elongate, paniculiform arrays, broadly so in robust plants (12.5–19 × 2.5–3 cm wide), consisting of short axillary and terminal racemiform clusters, proximal branches elongate in larger plants, branches glabrate to strigillose. |
(2–)30–500, in narrow to broad, thyrsiform to secund-pyramidal paniculiform arrays, branches recurved, secund, congested to lax. |
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Cypselae | narrowly obconic, 1.9–3.2 mm, sometimes with dark ridges, strigillose; pappi 1.9–5.2 mm (bristles sometimes clavate). |
0.7–2.7 mm, sparsely to densely strigillose; pappi 2.5–4.7 mm. |
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Solidago simplex |
Solidago velutina |
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Distribution |
AK; AZ; CO; ID; IN; MA; MD; ME; MI; MN; MT; ND; NH; NM; NY; OR; PA; SD; UT; VT; WA; WI; WV; WY; AB; BC; MB; NB; NT; ON; QC; SK; YT; Mexico
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AZ; CA; CO; ID; NM; NV; OR; TX; WY; Mexico
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Discussion | Varieties 7 (7 in the flora). The somewhat viscid-resinous heads of Solidago simplex are its most distinctive feature, separating it from similar sympatric species. G. S. Ringius (1985) did a detailed multivariate analysis of the S. spathulata/S. simplex complex (the latter under the name S. glutinosa). The cytogeography of the species complex was presented by Ringius and J. C. Semple (1987). Neither study included data on the next three species occurring in the southeastern United States. The species is divided into two subspecies and seven varieties following G. S Ringius (1985) and J. C. Semple et al. (1999). Three varieties occur in the diploid transcontinental subsp. simplex: var. simplex, var. nana, and var. chlorolepis. Four varieties occur in the eastern North American tetraploid-hexaploid subsp. randii: var. monticola, var. gillmanii, var. ontarioensis, and var. racemosa. Except for var. simplex, varieties are restricted to different habitats in relatively limited ranges. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 3 (2 in the flora). G. L. Nesom (1993b) merged Solidago californica, S. sparsiflora, and S. velutina without recognizing any infraspecific taxa, as did A. Cronquist (1994). J. C. Semple et al. (1990) compared S. californica and S. sparsiflora to S. nemoralis and found that all three are significantly different in a multivariate analysis. Evidence for separating the two subspecies of S. nemoralis was greater than the support for separating S. californica and S. sparsiflora. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 20, p. 113. | FNA vol. 20, p. 160. | ||||||||||||||||||||||||||||||||
Parent taxa | Asteraceae > tribe Astereae > Solidago > sect. Solidago > subsect. Humiles | Asteraceae > tribe Astereae > Solidago > sect. Solidago > subsect. Nemorales | ||||||||||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||||||||||
Subordinate taxa | ||||||||||||||||||||||||||||||||||
Synonyms | Aster velutinus | |||||||||||||||||||||||||||||||||
Name authority | Kunth: in A. von Humboldt et al., Nov. Gen. Sp. 4(fol.): 81. (1818) | de Candolle: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 5: 332. (1836) | ||||||||||||||||||||||||||||||||
Web links |