Solidago simplex |
Solidago ptarmicoides |
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alpine goldenrod, dune goldenrod, Mt. Albert goldenrod, Rand's goldenrod, spikelike goldenrod, sticky goldenrod |
prairie goldenrod, upland white aster, verge-d'or faux-ptarmica, white flat-top goldenrod |
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Habit | Plants 5–80 cm; caudices branching. | Plants 10–40 cm; caudices branching; vascular bundles and petiole bases marcescent (attached to old stems for more than a season). | ||||||||||||||||||||||||||||
Stems | 1–10+, ascending to erect, proximally glabrous, strigose in arrays. |
1–20+, erect, slender, glabrous proximally, hispid distally. |
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Leaves | basal and proximal cauline petiolate, blades narrowly oblanceolate, (20–)50–100(–160) × (2–)5–16(–56) mm, margins serrate to crenate, apices acute to obtuse, glabrous; mid and distal sessile, similar, blades lanceolate to linear, 12–45 × 2–19 mm, reduced distally, margins entire to sparsely serrate, sometimes resinous. |
basal and proximal cauline persistent, subpetiolate to sessile, blades sometimes 3-nerved, linear to linear-lanceolate, 60–100 × 7–10 mm, stiff, flat, margins entire or subentire, faces glabrous or sparsely hairy; mid to distal cauline sessile, blades linear oblanceolate to linear, 20–40 × 3–5 mm, reduced distally, margins entire. |
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Peduncles | 3.1–10.3 mm, glabrate to sparsely strigillose; bracteoles few, linear. |
29–36.5 mm, strigillose; bracteoles linear. |
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Involucres | campanulate, 3–7 mm. |
5–6 mm. |
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Ray florets | 7–16; laminae 2–5 × 0.7–0.9 mm. |
10–20, usually white, rarely pale yellow (conspicuous); laminae 7–7.3 × 1.4–1.6 mm. |
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Disc florets | 6–31; corollas 4–4.9 mm, lobes 0.6–1.3(–2) mm. |
30–36; corollas 3.8–4.1 mm, lobes 0.5–0.7 mm. |
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Phyllaries | (in 3–4 series) strongly unequal, often resinous; outer ovate, acute, inner linear-oblong, obtuse. |
in 4–5 series, usually prominently 1-nerved, linear, strongly unequal, outer acute, inner acute to obtuse, glabrous. |
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Heads | 3–150, not secund, in narrowly elongate, paniculiform arrays, broadly so in robust plants (12.5–19 × 2.5–3 cm wide), consisting of short axillary and terminal racemiform clusters, proximal branches elongate in larger plants, branches glabrate to strigillose. |
1–25(–50), in corymbiform arrays. |
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Cypselae | narrowly obconic, 1.9–3.2 mm, sometimes with dark ridges, strigillose; pappi 1.9–5.2 mm (bristles sometimes clavate). |
(obconic) 1–1.5 mm (ribbed), glabrous; pappi 3.4–4 mm (apically clavate). |
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2n | = 18. |
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Solidago simplex |
Solidago ptarmicoides |
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Phenology | Flowering Jul–Oct. | |||||||||||||||||||||||||||||
Habitat | Dry, sandy, usually calcareous soils, cracks in rocks, limestone pavements, rocky outcrops, grassy slopes, prairies | |||||||||||||||||||||||||||||
Elevation | 0–1500 m (0–4900 ft) | |||||||||||||||||||||||||||||
Distribution |
AK; AZ; CO; ID; IN; MA; MD; ME; MI; MN; MT; ND; NH; NM; NY; OR; PA; SD; UT; VT; WA; WI; WV; WY; AB; BC; MB; NB; NT; ON; QC; SK; YT; Mexico
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CO; CT; IA; IL; IN; MI; MN; MO; MT; NH; NY; OH; OK; SC; SD; VT; WI; WY; MB; ON; QC; SK
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Discussion | Varieties 7 (7 in the flora). The somewhat viscid-resinous heads of Solidago simplex are its most distinctive feature, separating it from similar sympatric species. G. S. Ringius (1985) did a detailed multivariate analysis of the S. spathulata/S. simplex complex (the latter under the name S. glutinosa). The cytogeography of the species complex was presented by Ringius and J. C. Semple (1987). Neither study included data on the next three species occurring in the southeastern United States. The species is divided into two subspecies and seven varieties following G. S Ringius (1985) and J. C. Semple et al. (1999). Three varieties occur in the diploid transcontinental subsp. simplex: var. simplex, var. nana, and var. chlorolepis. Four varieties occur in the eastern North American tetraploid-hexaploid subsp. randii: var. monticola, var. gillmanii, var. ontarioensis, and var. racemosa. Except for var. simplex, varieties are restricted to different habitats in relatively limited ranges. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
L. Brouillet and J. C. Semple (1981) summarized the morphologic, chemical, cytological, and hybridization data on Solidago ptarmicoides and concluded that, except for the white rays, it is a typical member of sect. Ptarmicoidei. The species hybridizes with S. rigida, S. ohioensis, S. riddellii, and perhaps other goldenrods. It is not known to hybridize with any species of aster (regardless of genus); the supposed Aster × Solidago hybrids are the basis of all reports of intergeneric hybrids in the literature. Horticultural hybrids involving S. ptarmicoides and other Solidago species have been treated as S. ×luteus (M. L. Green ex Dress) Brouillet & Semple (×Solidaster luteus M. L. Green ex Dress). Solidago ×bernardii B. Boivin [Oligoneuron ×bernardii (B. Boivin) G. L. Nesom] is the formal name applied to S. ptarmicoides × S. riddellii hybrids; those have cream rays. Solidago ×lutescens (Lindley ex de Candolle) B. Boivin [Diplopappus lutescens Lindley ex de Candolle; D. albus var. lutescens (Lindley ex de Candolle) Hooker ex Torrey & A. Gray; Aster lutescens (Lindley ex de Candolle) Hooker ex Torrey & A. Gray; A. ptarmicoides var. lutescens (Lindley ex de Candolle) A. Gray; Oligoneuron ×lutescens (Lindley ex de Candolle) G. L. Nesom] applies to hybrids between S. ptarmicoides and S. rigida or S. riddellii from the prairies; these also have cream colored rays. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 20, p. 113. | FNA vol. 20, p. 164. | ||||||||||||||||||||||||||||
Parent taxa | Asteraceae > tribe Astereae > Solidago > sect. Solidago > subsect. Humiles | Asteraceae > tribe Astereae > Solidago > sect. Ptarmicoidei | ||||||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||||||
Subordinate taxa | ||||||||||||||||||||||||||||||
Synonyms | Aster ptarmicoides, Diplopappus albus, Diplopappus ptarmicoides, Doellingeria ptarmicoides, Eucephalus albus, Heleastrum album, Inula alba, Oligoneuron album, S. asteroides, Unamia alba, Unamia ptarmicoides | |||||||||||||||||||||||||||||
Name authority | Kunth: in A. von Humboldt et al., Nov. Gen. Sp. 4(fol.): 81. (1818) | (Torrey & A. Gray) B. Boivin: Phytologia 23: 21. (1972) | ||||||||||||||||||||||||||||
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