Solidago simplex |
Solidago petiolaris |
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alpine goldenrod, dune goldenrod, Mt. Albert goldenrod, Rand's goldenrod, spikelike goldenrod, sticky goldenrod |
downy goldenrod, downy ragged goldenrod |
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Habit | Plants 5–80 cm; caudices branching. | Plants 40–150 cm; caudices stout, sometimes with long slender rhizomes. | ||||||||||||||||||||||||||||
Stems | 1–10+, ascending to erect, proximally glabrous, strigose in arrays. |
1–20(–50+), stout, finely puberulent or scabrous-puberulent at least distally. |
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Leaves | basal and proximal cauline petiolate, blades narrowly oblanceolate, (20–)50–100(–160) × (2–)5–16(–56) mm, margins serrate to crenate, apices acute to obtuse, glabrous; mid and distal sessile, similar, blades lanceolate to linear, 12–45 × 2–19 mm, reduced distally, margins entire to sparsely serrate, sometimes resinous. |
basal absent at flowering; cauline sessile to short-petiolate; blades usually lanceolate-elliptic or ovate, sometimes linear-lanceolate, 30–150 × 5–30 mm, thick and firm, margins entire or few toothed, somewhat to much reduced distally, abaxial faces sometimes resinous and shiny, glabrous or strigillose (hairs mostly 0.1–1.4 mm), adaxial glabrous or scabrous. |
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Peduncles | 3.1–10.3 mm, glabrate to sparsely strigillose; bracteoles few, linear. |
mostly 2–15 mm, bracteolate, sparsely to densely short hispid-strigose. |
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Involucres | campanulate, 3–7 mm. |
campanulate, 4.5–7.5 mm. |
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Ray florets | 7–16; laminae 2–5 × 0.7–0.9 mm. |
(5–)7–9; laminae 3–7 × 1–2 mm. |
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Disc florets | 6–31; corollas 4–4.9 mm, lobes 0.6–1.3(–2) mm. |
(8–)10–16; corollas 4–5 mm, lobes ca. 1–2 mm. |
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Phyllaries | (in 3–4 series) strongly unequal, often resinous; outer ovate, acute, inner linear-oblong, obtuse. |
in 3–4 series, unequal, linear-lanceolate, acute to attenuate, ± squarrose-tipped, glabrous or moderately strigose, sparsely to moderately stipitate-glandular, sometimes viscid. |
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Heads | 3–150, not secund, in narrowly elongate, paniculiform arrays, broadly so in robust plants (12.5–19 × 2.5–3 cm wide), consisting of short axillary and terminal racemiform clusters, proximal branches elongate in larger plants, branches glabrate to strigillose. |
10–190+ in paniculiform (rarely racemiform) arrays, usually elongate, usually leafy-bracteate, bracts similar to distal leaves but reduced; branches stiffly ascending, not secund, sometimes elongate. |
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Cypselae | narrowly obconic, 1.9–3.2 mm, sometimes with dark ridges, strigillose; pappi 1.9–5.2 mm (bristles sometimes clavate). |
3–4 mm, glabrous or glabrate; pappi ca. 4 mm. |
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2n | = 18, 36, 54. |
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Solidago simplex |
Solidago petiolaris |
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Phenology | Late Aug–Oct(–Nov). | |||||||||||||||||||||||||||||
Habitat | Woods and open places, especially sandy soils | |||||||||||||||||||||||||||||
Elevation | 0–1400[–2300] m (0–4600[–7500] ft) | |||||||||||||||||||||||||||||
Distribution |
AK; AZ; CO; ID; IN; MA; MD; ME; MI; MN; MT; ND; NH; NM; NY; OR; PA; SD; UT; VT; WA; WI; WV; WY; AB; BC; MB; NB; NT; ON; QC; SK; YT; Mexico
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AL; AR; FL; GA; IL; KS; LA; MO; NC; NE; NM; OK; SC; TX; Mexico (Coahuila)
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Discussion | Varieties 7 (7 in the flora). The somewhat viscid-resinous heads of Solidago simplex are its most distinctive feature, separating it from similar sympatric species. G. S. Ringius (1985) did a detailed multivariate analysis of the S. spathulata/S. simplex complex (the latter under the name S. glutinosa). The cytogeography of the species complex was presented by Ringius and J. C. Semple (1987). Neither study included data on the next three species occurring in the southeastern United States. The species is divided into two subspecies and seven varieties following G. S Ringius (1985) and J. C. Semple et al. (1999). Three varieties occur in the diploid transcontinental subsp. simplex: var. simplex, var. nana, and var. chlorolepis. Four varieties occur in the eastern North American tetraploid-hexaploid subsp. randii: var. monticola, var. gillmanii, var. ontarioensis, and var. racemosa. Except for var. simplex, varieties are restricted to different habitats in relatively limited ranges. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Solidago petiolaris is reported from Colorado but that report has not been confirmed. G. L. Nesom (1990j) discussed variation in the species and mapped the distribution. It is variable in leaf and phyllary shape and indument. Although several varieties have often been recognized in floras, the characters used to distinguish them form continua of variation that do not break into distinct groupings. Several general trends are worth noting. Plants in the Ozarks often have very resinous leaves; that does not appear to correlate with phyllary indument traits. Phyllary pubescence varies in a continuous fashion as well, with the numbers and distribution of hairs not breaking into discontinuous ranges. Diploids are known from throughout the range; one report of a tetraploid comes from North Carolina. Two reports of tetraploids from Florida were based on misidentified specimens. Plants with compact short arrays approach S. wrightii in this trait. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 20, p. 113. | FNA vol. 20, p. 118. | ||||||||||||||||||||||||||||
Parent taxa | Asteraceae > tribe Astereae > Solidago > sect. Solidago > subsect. Humiles | Asteraceae > tribe Astereae > Solidago > sect. Solidago > subsect. Thyrsiflorae | ||||||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||||||
Subordinate taxa | ||||||||||||||||||||||||||||||
Synonyms | Aster lindheimeranus, Aster petiolaris, S. angusta, S. harperi, S. lindheimeriana, S. milleriana, S. petiolaris var. angusta, S. petiolaris var. squarrulosa, S. petiolaris var. wardii, S. squarrulosa, S. wardii | |||||||||||||||||||||||||||||
Name authority | Kunth: in A. von Humboldt et al., Nov. Gen. Sp. 4(fol.): 81. (1818) | Aiton: Hort. Kew. 3: 216. (1789) | ||||||||||||||||||||||||||||
Web links |