Solidago simplex
Solidago hispida
alpine goldenrod, dune goldenrod, Mt. Albert goldenrod, Rand's goldenrod, spikelike goldenrod, sticky goldenrod
hairy goldenrod, verge d'or hispide
1–10+, ascending to erect, proximally glabrous, strigose in arrays.
1(–5), erect, simple, glabrous or moderately to densely hispido-villous, sometimes hairs appressed.
basal and proximal cauline petiolate, blades narrowly oblanceolate, (20–)50–100(–160) × (2–)5–16(–56) mm, margins serrate to crenate, apices acute to obtuse, glabrous;
mid and distal sessile, similar, blades lanceolate to linear, 12–45 × 2–19 mm, reduced distally, margins entire to sparsely serrate, sometimes resinous.
basal and proximal cauline tapering to winged petioles, blades broadly oblanceolate to obovate or elliptic, 35–200 (including petioles) × 15–60 mm, margins serrate or crenate, apices acute, faces sparsely to densely hispido-villous, rarely glabrous;
mid and distal cauline sessile, blades elliptic, 15–30 × 5–7 mm, rapidly reduced distally, margins entire.
3.1–10.3 mm, glabrate to sparsely strigillose;
bracteoles few, linear.
1.5–2.5 mm, hispido-villous to canescent.
campanulate, 3–7 mm.
campanulate, 4–6 mm.
7–16;
laminae 2–5 × 0.7–0.9 mm.
6–14;
laminae 1.5–4.5 × 0.5–1 mm.
6–31;
corollas 4–4.9 mm, lobes 0.6–1.3(–2) mm.
6–12;
corollas 3–4.5 mm, lobes 0.6–1.2 mm.
(in 3–4 series) strongly unequal, often resinous;
outer ovate, acute, inner linear-oblong, obtuse.
in 3–4 series, appressed, strongly unequal, midribs and tips conspicuously green, margins white, scarious, apices obtuse, glabrous or moderately strigose.
3–150, not secund, in narrowly elongate, paniculiform arrays, broadly so in robust plants (12.5–19 × 2.5–3 cm wide), consisting of short axillary and terminal racemiform clusters, proximal branches elongate in larger plants, branches glabrate to strigillose.
8–250+ in usually wand-paniculiform arrays of short axillary and terminal racemiform, non-secund clusters, sometimes proximal branches elongated, ascending and bearing short axillary and terminal racemiform clusters.
narrowly obconic, 1.9–3.2 mm, sometimes with dark ridges, strigillose;
pappi 1.9–5.2 mm (bristles sometimes clavate).
(narrowly obconic) 1–2.5 mm, glabrous;
pappi 2.5–4 mm (sometimes distinctly clavate).
= 18.
Solidago simplex
Solidago hispida
Varieties 7 (7 in the flora).
The somewhat viscid-resinous heads of Solidago simplex are its most distinctive feature, separating it from similar sympatric species. G. S. Ringius (1985) did a detailed multivariate analysis of the S. spathulata/S. simplex complex (the latter under the name S. glutinosa). The cytogeography of the species complex was presented by Ringius and J. C. Semple (1987). Neither study included data on the next three species occurring in the southeastern United States.
The species is divided into two subspecies and seven varieties following G. S Ringius (1985) and J. C. Semple et al. (1999). Three varieties occur in the diploid transcontinental subsp. simplex: var. simplex, var. nana, and var. chlorolepis. Four varieties occur in the eastern North American tetraploid-hexaploid subsp. randii: var. monticola, var. gillmanii, var. ontarioensis, and var. racemosa. Except for var. simplex, varieties are restricted to different habitats in relatively limited ranges.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Ray floret color can fade with time, both in the field and on dried specimens, making it difficult to distinguish Solidago hispida from S. bicolor, in which it has sometimes been included. The latter tends to have slightly broader phyllaries and more obviously clavate pappus bristles. The two co-occur in some populations but do not appear to form hybrid swarms with a range in ray corolla color. Three varieties have been described on the basis of growth and pubescence features and are sometimes recognized. These may represent only extremes in continua of variation. Variety lanata has densely villous stems and leaves; such plants occur scattered through much of the range of the species. Two glabrous-stemmed varieties have been described; plants of var. tonsa are relatively small and were first described from Newfoundland; var. huronensis includes relatively tall glabrous plants found growing in sand dunes along the shore of Lake Huron in Michigan and Ontario. The latter plants have been treated as var. tonsa but the name is misapplied. Other varietal names are based on minor variations.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Basal and cauline leaves spatulate to orbiculate, apices obtuse to rounded; cypselae moderately to densely strigose (S. simplex subsp. simplex) | → 2 |
1. Basal and cauline leaves spatulate to obovate, apices usually acute; cypselae sparsely to moderately strigose (S. simplex subsp. randii) | → 5 |
2. Cauline leaves 5–16 (longest 60+ mm); w North America, disjunct along shores of upper Great Lakes and in s Quebec | var. simplex |
2. Cauline leaves 2–7 (longest to 60 mm); alpine slopes, w North America, Mt. Albert, Gaspé, Quebec | → 3 |
3. Shortest phyllaries 1.4–2 mm; disc corollas 3.9–4.2 mm; serpentine soils, Mt. Albert, Gaspé, Quebec | var. chlorolepis |
3. Shortest phyllaries 2.1–3.1 mm; disc corollas 4.2–5.4 mm; alpine slopes, Rocky Mountains, Cascade Mountains, Vancouver Island. | → 4 |
4. Basal leaves usually spatulate, rarely broadly elliptic, proximal blades gradually attenuated to winged petioles, margins weakly crenate; alpine slopes, Rocky Mountains | var. simplex |
4. Basal leaves orbiculate to broadly elliptic, proximal blades abruptly attenuated to winged petioles, margins distinctly crenate; alpine slopes of Cascade Mountains, Vancouver Island | var. nana |
5. Proximal cauline leaves spatulate to obovate, (5–)10–42 mm wide; leaf margins often sharply serrate | → 6 |
5. Proximal cauline leaves lanceolate to narrowly spatulate, 2–10(–21.5) mm wide; leaf margins entire or dentate, rarely serrate | → 7 |
6. Plants (10–)18–53(–83) cm, arrays tending to be compact, virgate; proximal cauline leaves (5–)10–22(–31) mm wide, margins crenate to serrate; barren, rocky, non-alpine uplands; se Quebec, New England, New York | var. monticola |
6. Plants (20–)38–73(–84) cm, robust, arrays tending to be paniculiform; proximal cauline leaves (6–)7.5–24(–42) mm wide, margins often sharply serrate; sand dunes, Lake Michigan, n Lake Huron | var. gillmanii |
7. Cauline leaves (3–)12–33(–58); peduncle bracteoles 2+; calcareous rocky riverbanks, e Quebec s to West Virginia and Maryland | var. racemosa |
7. Cauline leaves (2–)4–13(–24); peduncle bracteoles 1–3(–4); rocky shores, Lake Superior, n Lake Huron | var. ontarioensis |
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