Solidago simplex |
Solidago arenicola |
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alpine goldenrod, dune goldenrod, Mt. Albert goldenrod, Rand's goldenrod, spikelike goldenrod, sticky goldenrod |
sand goldenrod, southern racemose goldenrod |
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Habit | Plants 5–80 cm; caudices branching. | Plants 35–80 cm; rhizomes horizontal to ascending, short, this and its branches apically caudexlike, to 5 mm thick, mostly embedded in imbricate chaffy bases of older leaves. | ||||||||||||||||||||||||||||
Stems | 1–10+, ascending to erect, proximally glabrous, strigose in arrays. |
1–3, terete, shallowly multicostate, glabrous, sparsely to moderately hirtellous distally (arrays). |
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Leaves | basal and proximal cauline petiolate, blades narrowly oblanceolate, (20–)50–100(–160) × (2–)5–16(–56) mm, margins serrate to crenate, apices acute to obtuse, glabrous; mid and distal sessile, similar, blades lanceolate to linear, 12–45 × 2–19 mm, reduced distally, margins entire to sparsely serrate, sometimes resinous. |
basal and proximal cauline winged-petiolate; blades mostly spatulate to oblanceolate, 100–150 × 15–32 mm, tapering, margins distally coarsely but shallowly serrate, apices acute to narrowly rounded, often narrowed to short, often brown callused tips; mid to distal cauline progressively more narrowly oblanceolate and acute, then to elliptic or lanceolate-elliptic, 35–60 × 6–10 mm, reduced distally, grading into widely ascending to spreading or reflexed bracts; faces abaxially pale, midnerves raised, deep green, level reticulum of branch nerves, margins entire, ciliate, adaxially deep green with only impressed midnerve evident. |
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Peduncles | 3.1–10.3 mm, glabrate to sparsely strigillose; bracteoles few, linear. |
stiffly, widely ascending, angulate, slightly compressed, stubby-bracteolate; bracts grading into phyllaries, sparsely hirtellous. |
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Involucres | campanulate, 3–7 mm. |
turbinate, 8–12 mm. |
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Ray florets | 7–16; laminae 2–5 × 0.7–0.9 mm. |
6–10; laminae 3–4 mm. |
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Disc florets | 6–31; corollas 4–4.9 mm, lobes 0.6–1.3(–2) mm. |
11–14; corollas 6 mm, lobes ca. 2 mm. |
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Phyllaries | (in 3–4 series) strongly unequal, often resinous; outer ovate, acute, inner linear-oblong, obtuse. |
(in 3–4 series) 1–1.3 mm wide, unequal, pilose-ciliate; outer mostly green, oblong, blunt, inner spatulate or linear-oblanceolate. |
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Heads | 3–150, not secund, in narrowly elongate, paniculiform arrays, broadly so in robust plants (12.5–19 × 2.5–3 cm wide), consisting of short axillary and terminal racemiform clusters, proximal branches elongate in larger plants, branches glabrate to strigillose. |
10–50 (1–5+ per branch), in racemiform or narrowly paniculiform arrays, mostly with primary branches widely to narrowly ascending, proximalmost usually longest. |
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Cypselae | narrowly obconic, 1.9–3.2 mm, sometimes with dark ridges, strigillose; pappi 1.9–5.2 mm (bristles sometimes clavate). |
cylindric to somewhat compressed-obconic, 1.5–4 mm, typically 5-ribbed, glabrous, abruptly narrowed to short “neck” surmounted by narrow, brownish pappus disc; pappi (white) 3–4 mm. |
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2n | = 36. |
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Solidago simplex |
Solidago arenicola |
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Phenology | Flowering (Aug–)Sep–Oct. | |||||||||||||||||||||||||||||
Habitat | Mesic woods in deep sandy alluvium | |||||||||||||||||||||||||||||
Elevation | 100–200 m (300–700 ft) | |||||||||||||||||||||||||||||
Distribution |
AK; AZ; CO; ID; IN; MA; MD; ME; MI; MN; MT; ND; NH; NM; NY; OR; PA; SD; UT; VT; WA; WI; WV; WY; AB; BC; MB; NB; NT; ON; QC; SK; YT; Mexico
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AL; TN |
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Discussion | Varieties 7 (7 in the flora). The somewhat viscid-resinous heads of Solidago simplex are its most distinctive feature, separating it from similar sympatric species. G. S. Ringius (1985) did a detailed multivariate analysis of the S. spathulata/S. simplex complex (the latter under the name S. glutinosa). The cytogeography of the species complex was presented by Ringius and J. C. Semple (1987). Neither study included data on the next three species occurring in the southeastern United States. The species is divided into two subspecies and seven varieties following G. S Ringius (1985) and J. C. Semple et al. (1999). Three varieties occur in the diploid transcontinental subsp. simplex: var. simplex, var. nana, and var. chlorolepis. Four varieties occur in the eastern North American tetraploid-hexaploid subsp. randii: var. monticola, var. gillmanii, var. ontarioensis, and var. racemosa. Except for var. simplex, varieties are restricted to different habitats in relatively limited ranges. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Solidago arenicola is known from the flood plain of the Locust Fork River, Blount County, Alabama. The Tennessee plants that have been treated as Solidago simplex var. racemosa are included here in this large-headed southern species. Further work is needed to confirm this placement. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 20, p. 113. | FNA vol. 20, p. 116. | ||||||||||||||||||||||||||||
Parent taxa | Asteraceae > tribe Astereae > Solidago > sect. Solidago > subsect. Humiles | Asteraceae > tribe Astereae > Solidago > sect. Solidago > subsect. Humiles | ||||||||||||||||||||||||||||
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Name authority | Kunth: in A. von Humboldt et al., Nov. Gen. Sp. 4(fol.): 81. (1818) | B. R. Keener & Kral: Sida 20: 1589, fig. 1. (2003) | ||||||||||||||||||||||||||||
Web links |