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alpine goldenrod, dune goldenrod, Mt. Albert goldenrod, Rand's goldenrod, spikelike goldenrod, sticky goldenrod

sand goldenrod, southern racemose goldenrod

Habit Plants 5–80 cm; caudices branching. Plants 35–80 cm; rhizomes horizontal to ascending, short, this and its branches apically caudexlike, to 5 mm thick, mostly embedded in imbricate chaffy bases of older leaves.
Stems

1–10+, ascending to erect, proximally glabrous, strigose in arrays.

1–3, terete, shallowly multicostate, glabrous, sparsely to moderately hirtellous distally (arrays).

Leaves

basal and proximal cauline petiolate, blades narrowly oblanceolate, (20–)50–100(–160) × (2–)5–16(–56) mm, margins serrate to crenate, apices acute to obtuse, glabrous;

mid and distal sessile, similar, blades lanceolate to linear, 12–45 × 2–19 mm, reduced distally, margins entire to sparsely serrate, sometimes resinous.

basal and proximal cauline winged-petiolate;

blades mostly spatulate to oblanceolate, 100–150 × 15–32 mm, tapering, margins distally coarsely but shallowly serrate, apices acute to narrowly rounded, often narrowed to short, often brown callused tips;

mid to distal cauline progressively more narrowly oblanceolate and acute, then to elliptic or lanceolate-elliptic, 35–60 × 6–10 mm, reduced distally, grading into widely ascending to spreading or reflexed bracts;

faces abaxially pale, midnerves raised, deep green, level reticulum of branch nerves, margins entire, ciliate, adaxially deep green with only impressed midnerve evident.

Peduncles

3.1–10.3 mm, glabrate to sparsely strigillose;

bracteoles few, linear.

stiffly, widely ascending, angulate, slightly compressed, stubby-bracteolate;

bracts grading into phyllaries, sparsely hirtellous.

Involucres

campanulate, 3–7 mm.

turbinate, 8–12 mm.

Ray florets

7–16;

laminae 2–5 × 0.7–0.9 mm.

6–10;

laminae 3–4 mm.

Disc florets

6–31;

corollas 4–4.9 mm, lobes 0.6–1.3(–2) mm.

11–14;

corollas 6 mm, lobes ca. 2 mm.

Phyllaries

(in 3–4 series) strongly unequal, often resinous;

outer ovate, acute, inner linear-oblong, obtuse.

(in 3–4 series) 1–1.3 mm wide, unequal, pilose-ciliate;

outer mostly green, oblong, blunt, inner spatulate or linear-oblanceolate.

Heads

3–150, not secund, in narrowly elongate, paniculiform arrays, broadly so in robust plants (12.5–19 × 2.5–3 cm wide), consisting of short axillary and terminal racemiform clusters, proximal branches elongate in larger plants, branches glabrate to strigillose.

10–50 (1–5+ per branch), in racemiform or narrowly paniculiform arrays, mostly with primary branches widely to narrowly ascending, proximalmost usually longest.

Cypselae

narrowly obconic, 1.9–3.2 mm, sometimes with dark ridges, strigillose;

pappi 1.9–5.2 mm (bristles sometimes clavate).

cylindric to somewhat compressed-obconic, 1.5–4 mm, typically 5-ribbed, glabrous, abruptly narrowed to short “neck” surmounted by narrow, brownish pappus disc;

pappi (white) 3–4 mm.

2n

= 36.

Solidago simplex

Solidago arenicola

Phenology Flowering (Aug–)Sep–Oct.
Habitat Mesic woods in deep sandy alluvium
Elevation 100–200 m (300–700 ft)
Distribution
from FNA
AK; AZ; CO; ID; IN; MA; MD; ME; MI; MN; MT; ND; NH; NM; NY; OR; PA; SD; UT; VT; WA; WI; WV; WY; AB; BC; MB; NB; NT; ON; QC; SK; YT; Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; TN
[BONAP county map]
Discussion

Varieties 7 (7 in the flora).

The somewhat viscid-resinous heads of Solidago simplex are its most distinctive feature, separating it from similar sympatric species. G. S. Ringius (1985) did a detailed multivariate analysis of the S. spathulata/S. simplex complex (the latter under the name S. glutinosa). The cytogeography of the species complex was presented by Ringius and J. C. Semple (1987). Neither study included data on the next three species occurring in the southeastern United States.

The species is divided into two subspecies and seven varieties following G. S Ringius (1985) and J. C. Semple et al. (1999). Three varieties occur in the diploid transcontinental subsp. simplex: var. simplex, var. nana, and var. chlorolepis. Four varieties occur in the eastern North American tetraploid-hexaploid subsp. randii: var. monticola, var. gillmanii, var. ontarioensis, and var. racemosa. Except for var. simplex, varieties are restricted to different habitats in relatively limited ranges.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Solidago arenicola is known from the flood plain of the Locust Fork River, Blount County, Alabama. The Tennessee plants that have been treated as Solidago simplex var. racemosa are included here in this large-headed southern species. Further work is needed to confirm this placement.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal and cauline leaves spatulate to orbiculate, apices obtuse to rounded; cypselae moderately to densely strigose (S. simplex subsp. simplex)
→ 2
1. Basal and cauline leaves spatulate to obovate, apices usually acute; cypselae sparsely to moderately strigose (S. simplex subsp. randii)
→ 5
2. Cauline leaves 5–16 (longest 60+ mm); w North America, disjunct along shores of upper Great Lakes and in s Quebec
var. simplex
2. Cauline leaves 2–7 (longest to 60 mm); alpine slopes, w North America, Mt. Albert, Gaspé, Quebec
→ 3
3. Shortest phyllaries 1.4–2 mm; disc corollas 3.9–4.2 mm; serpentine soils, Mt. Albert, Gaspé, Quebec
var. chlorolepis
3. Shortest phyllaries 2.1–3.1 mm; disc corollas 4.2–5.4 mm; alpine slopes, Rocky Mountains, Cascade Mountains, Vancouver Island.
→ 4
4. Basal leaves usually spatulate, rarely broadly elliptic, proximal blades gradually attenuated to winged petioles, margins weakly crenate; alpine slopes, Rocky Mountains
var. simplex
4. Basal leaves orbiculate to broadly elliptic, proximal blades abruptly attenuated to winged petioles, margins distinctly crenate; alpine slopes of Cascade Mountains, Vancouver Island
var. nana
5. Proximal cauline leaves spatulate to obovate, (5–)10–42 mm wide; leaf margins often sharply serrate
→ 6
5. Proximal cauline leaves lanceolate to narrowly spatulate, 2–10(–21.5) mm wide; leaf margins entire or dentate, rarely serrate
→ 7
6. Plants (10–)18–53(–83) cm, arrays tending to be compact, virgate; proximal cauline leaves (5–)10–22(–31) mm wide, margins crenate to serrate; barren, rocky, non-alpine uplands; se Quebec, New England, New York
var. monticola
6. Plants (20–)38–73(–84) cm, robust, arrays tending to be paniculiform; proximal cauline leaves (6–)7.5–24(–42) mm wide, margins often sharply serrate; sand dunes, Lake Michigan, n Lake Huron
var. gillmanii
7. Cauline leaves (3–)12–33(–58); peduncle bracteoles 2+; calcareous rocky riverbanks, e Quebec s to West Virginia and Maryland
var. racemosa
7. Cauline leaves (2–)4–13(–24); peduncle bracteoles 1–3(–4); rocky shores, Lake Superior, n Lake Huron
var. ontarioensis
Source FNA vol. 20, p. 113. FNA vol. 20, p. 116.
Parent taxa Asteraceae > tribe Astereae > Solidago > sect. Solidago > subsect. Humiles Asteraceae > tribe Astereae > Solidago > sect. Solidago > subsect. Humiles
Sibling taxa
S. albopilosa, S. altiplanities, S. altissima, S. arenicola, S. arguta, S. auriculata, S. bicolor, S. brachyphylla, S. buckleyi, S. caesia, S. canadensis, S. confinis, S. curtisii, S. delicatula, S. drummondii, S. elongata, S. erecta, S. faucibus, S. fistulosa, S. flexicaulis, S. gattingeri, S. gigantea, S. glomerata, S. guiradonis, S. hispida, S. houghtonii, S. juliae, S. juncea, S. kralii, S. lancifolia, S. latissimifolia, S. leavenworthii, S. leiocarpa, S. lepida, S. ludoviciana, S. macrophylla, S. missouriensis, S. mollis, S. multiradiata, S. nana, S. nemoralis, S. nitida, S. odora, S. ohioensis, S. ouachitensis, S. patula, S. petiolaris, S. pinetorum, S. plumosa, S. ptarmicoides, S. puberula, S. pulchra, S. radula, S. riddellii, S. rigida, S. roanensis, S. rugosa, S. rupestris, S. sciaphila, S. sempervirens, S. shortii, S. spathulata, S. speciosa, S. spectabilis, S. sphacelata, S. spithamaea, S. squarrosa, S. stricta, S. tarda, S. tortifolia, S. uliginosa, S. ulmifolia, S. velutina, S. verna, S. villosicarpa, S. wrightii
S. albopilosa, S. altiplanities, S. altissima, S. arguta, S. auriculata, S. bicolor, S. brachyphylla, S. buckleyi, S. caesia, S. canadensis, S. confinis, S. curtisii, S. delicatula, S. drummondii, S. elongata, S. erecta, S. faucibus, S. fistulosa, S. flexicaulis, S. gattingeri, S. gigantea, S. glomerata, S. guiradonis, S. hispida, S. houghtonii, S. juliae, S. juncea, S. kralii, S. lancifolia, S. latissimifolia, S. leavenworthii, S. leiocarpa, S. lepida, S. ludoviciana, S. macrophylla, S. missouriensis, S. mollis, S. multiradiata, S. nana, S. nemoralis, S. nitida, S. odora, S. ohioensis, S. ouachitensis, S. patula, S. petiolaris, S. pinetorum, S. plumosa, S. ptarmicoides, S. puberula, S. pulchra, S. radula, S. riddellii, S. rigida, S. roanensis, S. rugosa, S. rupestris, S. sciaphila, S. sempervirens, S. shortii, S. simplex, S. spathulata, S. speciosa, S. spectabilis, S. sphacelata, S. spithamaea, S. squarrosa, S. stricta, S. tarda, S. tortifolia, S. uliginosa, S. ulmifolia, S. velutina, S. verna, S. villosicarpa, S. wrightii
Subordinate taxa
S. simplex var. chlorolepis, S. simplex var. gillmanii, S. simplex var. monticola, S. simplex var. nana, S. simplex var. ontarioensis, S. simplex var. racemosa, S. simplex var. simplex
Name authority Kunth: in A. von Humboldt et al., Nov. Gen. Sp. 4(fol.): 81. (1818) B. R. Keener & Kral: Sida 20: 1589, fig. 1. (2003)
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