Solidago radula |
Solidago villosicarpa |
|
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rough goldenrod, western rough goldenrod |
glandular wand goldenrod, hairy-seed goldenrod |
|
Habit | Plants 30–90 cm; caudices, sometimes also creeping rhizomes as well. | Plants loosely cespitose, 45–150 cm; caudices short, woody, or short rhizomes. |
Stems | usually 1–3, ascending to erect, scabrous to loosely puberulent. |
usually single, erect (proximally medium to dark brown distally lighter, sometimes cyanotic, usually rounded, shallowly many ribbed), sparsely finely hispido-strigose proximally to densely so in arrays. |
Leaves | basal and proximal usually withering by flowering, tapering to long-winged petioles, blades oblanceolate, 30–100 × 7–20(–30) mm, margins serrate or crenate, mid usually largest, apices acute to obtuse, acuminate, faces scabrous; mid and distal cauline subsessile (1 mm) or sessile, blades (sometimes ± shiny) elliptic to oblanceolate, 10–50 × 5–15(–25) mm, greatly reduced distally, grading into bracts, firm, bases convex-cuneate to rounded, margins finely serrate, often 3-nerved, nerves usually distinct abaxially, faces distinctly scabrous. |
rosettes often present at flowering, early leaves smaller; basal and proximal cauline gradually to abruptly tapering to winged petioles, blades elliptic to ovate-elliptic, 90–210 (including petioles) × 40–70 mm, margins serrate-serrulate, ciliate, apices obtuse to broadly acute, faces abaxially glabrous or sparsely strigose, adaxially glabrous or sparsely strigose, mostly on nerves; mid to distal cauline sessile, blades lanceolate or elliptic, 20–650 × 10–30 mm, gradually reduced distally, becoming entire, apices acute to acuminate. |
Peduncles | 0.5–2 mm; bracteoles 1–5, linear-lanceolate to ovate, minute, grading into phyllaries distally. |
0.5–11 mm, densely short hispido-strigose; bracteoles 5–10+, grading into phyllaries, sparsely short hispido-strigose and sparsely to moderately glandular. |
Involucres | narrowly campanulate, 3–5 mm. |
campanulate, 5–8 mm. |
Ray florets | 4–7; laminae 2–3.5 × 0.2–0.7 mm. |
4–8; laminae 5–7 × 1–2 mm. |
Disc florets | 4–6; corollas 3 mm, lobes 1 mm. |
10–18; corollas 5–7 mm, lobes 1.5–2.2 mm. |
Phyllaries | in 3–4 series, unequal, oblong, midnerves swollen distally, obtuse or acute to slightly acuminate. |
in 4–5 series, appressed, strongly unequal, outer ovate, acute, mid and inner broadly oblong (appressed), obtuse or rounded, sparsely strigose and moderately finely stipitate-glandular. |
Heads | 20–260, in paniculiform arrays, narrowly to broadly secund, pyramidal, branches recurved, secund. |
50–100+ (1–10 per branch, more on much elongated proximal branches), in elongate to thyrsiform-paniculiform arrays 7–22 × 3–6 cm; branches ascending, racemiform or paniculiform, not secund, longest to 1/2 length of arrays. |
Cypselae | 1.5–2.5 mm, sparsely to moderately short-strigose; pappi 3 mm. |
(fusiform to obconic) 2.5–3 mm, moderately long-strigose; pappi 4–6 mm (some clavate). |
2n | = 18, 36. |
|
Solidago radula |
Solidago villosicarpa |
|
Phenology | Flowering Aug–Oct. | Flowering Sep. |
Habitat | Open rocky places, dry woods, especially calcareous soils | Sandy soils, live oak scrub on dunes, roadsides, open pine-oak woods, Atlantic coastal plain |
Elevation | 0–600 m (0–2000 ft) | 10–20 m (0–100 ft) |
Distribution |
AR; GA; IL; KS; KY; LA; MO; NC; OK; SC; TX
|
NC |
Discussion | Solidago radula is disjunct in Georgia, North Carolina, and South Carolina. J. R. Beaudry (1969) reported a diploid from Smithville, Dekalb County, Tennessee; that has not been confirmed. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Solidago villosicarpa is known only from New Hanover, Onslow, and Pender counties. It has a habit similar to those of S. hispida and S. squarrosa, leaves similar to those of S. erecta, and stem pubescence similar to that of S. puberula. The large, glandular involucres are unique within subsect. Squarrosae. It was compared to S. sciaphila when first described; it is not very similar to that Midwestern species. It is likely either a large-headed diploid like S. squarrosa or possibly an allopolyploid. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 20, p. 162. | FNA vol. 20, p. 122. |
Parent taxa | Asteraceae > tribe Astereae > Solidago > sect. Solidago > subsect. Nemorales | Asteraceae > tribe Astereae > Solidago > sect. Solidago > subsect. Squarrosae |
Sibling taxa | ||
Synonyms | Aster decemflora, S. decemflora, S. laeta, S. pendula, S. radula var. laeta, S. radula var. rotundifolia, S. radula var. stenolepis, S. rotundifolia, S. scaberrima | |
Name authority | Nuttall: J. Acad. Nat. Sci. Philadelphia 7: 102. (1834) | LeBlond: Sida 19: 292, figs. 1–6. (2000) |
Web links |