Solidago radula |
Solidago missouriensis |
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rough goldenrod, western rough goldenrod |
Missouri goldenrod, prairie goldenrod |
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Habit | Plants 30–90 cm; caudices, sometimes also creeping rhizomes as well. | Plants (10–)30–80 cm; rhizomes short to long. |
Stems | usually 1–3, ascending to erect, scabrous to loosely puberulent. |
1–50+, erect, glabrous or sometimes sparsely strigose in arrays; fascicles of small lateral branch leaves often present in axils. |
Leaves | basal and proximal usually withering by flowering, tapering to long-winged petioles, blades oblanceolate, 30–100 × 7–20(–30) mm, margins serrate or crenate, mid usually largest, apices acute to obtuse, acuminate, faces scabrous; mid and distal cauline subsessile (1 mm) or sessile, blades (sometimes ± shiny) elliptic to oblanceolate, 10–50 × 5–15(–25) mm, greatly reduced distally, grading into bracts, firm, bases convex-cuneate to rounded, margins finely serrate, often 3-nerved, nerves usually distinct abaxially, faces distinctly scabrous. |
proximal cauline tapering to long, winged petioles, blades oblanceolate to linear-oblanceolate, 50–100(–200) (including petiole) × (5–)10–20(–30) mm, margins entire to serrulate, usually 3-nerved (2 larger lateral nerves), apices acute, mucronate to acuminate and somewhat spinulose, glabrous; mid to distal cauline sessile, blades lanceolate to linear, 40–60 × (2–)4–14 mm, rapidly reduced distally, margins entire, ciliate, faces glabrous. |
Peduncles | 0.5–2 mm; bracteoles 1–5, linear-lanceolate to ovate, minute, grading into phyllaries distally. |
1.4–5 mm, glabrous or sparsely strigose; bracteoles 0–3+, linear to lanceolate. |
Involucres | narrowly campanulate, 3–5 mm. |
narrowly to broadly campanulate, 2.5–4.5 mm. |
Ray florets | 4–7; laminae 2–3.5 × 0.2–0.7 mm. |
5–14; laminae 1.5–2(–4) × 0.2–0.5(–0.75) mm. |
Disc florets | 4–6; corollas 3 mm, lobes 1 mm. |
(6–)8–20; corollas (2–)3–4 mm, lobes 0.4–1 mm. |
Phyllaries | in 3–4 series, unequal, oblong, midnerves swollen distally, obtuse or acute to slightly acuminate. |
in 3–4 series, strongly unequal, margins ciliate-fimbriate, especially apically; outer ovate to lanceolate, acute to rounded, inner linear-ovate to oblong or linear-lanceolate, obtuse to rounded. |
Heads | 20–260, in paniculiform arrays, narrowly to broadly secund, pyramidal, branches recurved, secund. |
10–210 in paniculiform arrays, broadly secund-pyramidal or more rhombic to transversely rhombic, (1.5–)3–12(–20) × (1.5–)3–12 cm; branches glabrous with secund heads spreading and arching, sometimes ascending with non-secund heads. |
Cypselae | 1.5–2.5 mm, sparsely to moderately short-strigose; pappi 3 mm. |
(obconic) 1–2 mm, glabrous or sparsely strigose; pappi 2.5–3 mm. |
2n | = 18, 36. |
= 18, 36. |
Solidago radula |
Solidago missouriensis |
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Phenology | Flowering Aug–Oct. | Flowering (Jul–)Aug–Oct. |
Habitat | Open rocky places, dry woods, especially calcareous soils | Open sandy and rocky soils, clay soils, prairies, grasslands, pastures, open conifers forests in foothills and proximal elevations of mountains, sandstone ledges, limestone glades, disturbed soils, roadsides |
Elevation | 0–600 m (0–2000 ft) | 200–2200+ m (700–7200+ ft) |
Distribution |
AR; GA; IL; KS; KY; LA; MO; NC; OK; SC; TX
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AR; AZ; CO; ID; IL; IN; KS; KY; MI; MN; MO; MT; ND; NE; NM; NV; OK; OR; SD; TN; TX; UT; WA; WY; AB; BC; MB; ON; SK; Mexico (Coahuila)
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Discussion | Solidago radula is disjunct in Georgia, North Carolina, and South Carolina. J. R. Beaudry (1969) reported a diploid from Smithville, Dekalb County, Tennessee; that has not been confirmed. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Solidago missouriensis was often introduced along railroad lines farther east. It is a highly variable species. In the east, it can be similar to S. juncea and is not always easily distinguished where ranges overlap. In the west, it can similar to smaller plants of S. spectabilis. It is distinguished from the related species by its usually 3-nerved proximal leaves and the usually thin, elongate rhizomes. Across the prairies the species is known to be diploid only (2n = 18). In the Rocky Mountains, tetraploids (2n = 36) are common, the diploids infrequent. A number of varieties have been described. Shorter, often larger-headed plants (tetraploids when known) from the Rocky Mountains have been treated as var. missouriensis (including var. extraria). Taller, more leafy-stemmed plants, mostly from the eastern half of the range, but occasionally west to Washington, have been treated as var. fasciculata. Plants from Arizona, Colorado, and New Mexico with long, linear leaves have been treated as var. tenuissima. Larger-headed plants with narrow bracts from prairies west of the Cascades in Oregon and Washington have been treated as var. tolmieana. A. Cronquist (1994) opted not to recognize varieties, noting that all appeared to grade continuously into each other. A detailed study of the species is needed. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 20, p. 162. | FNA vol. 20, p. 142. |
Parent taxa | Asteraceae > tribe Astereae > Solidago > sect. Solidago > subsect. Nemorales | Asteraceae > tribe Astereae > Solidago > sect. Solidago > subsect. Junceae |
Sibling taxa | ||
Synonyms | Aster decemflora, S. decemflora, S. laeta, S. pendula, S. radula var. laeta, S. radula var. rotundifolia, S. radula var. stenolepis, S. rotundifolia, S. scaberrima | Aster marshallii, Aster missouriensis, Aster tolmieanus, Doria concinna, Doria glaberrima, Doria glaberrima var. montana, S. concinna, S. duriuscula, S. glaberrima, S. glaberrima var. montana, S. glaberrima var. moritura, S. glaucophylla, S. hapemaniana, S. marshallii, S. missouriensis var. extraria, S. missouriensis var. fasciculata, S. missouriensis var. glaberrima, S. missouriensis var. montana, S. missouriensis var. tenuissima, S. missouriensis var. tolmieana, S. moritura, S. tenuissima, S. tolmieana |
Name authority | Nuttall: J. Acad. Nat. Sci. Philadelphia 7: 102. (1834) | Nuttall: J. Acad. Nat. Sci. Philadelphia 7: 32. (1834) |
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