Solidago radula |
Solidago houghtonii |
|
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rough goldenrod, western rough goldenrod |
Houghton's goldenrod |
|
Habit | Plants 30–90 cm; caudices, sometimes also creeping rhizomes as well. | Plants 30–60+ cm; caudices branching; vascular bundles and petiole bases marcescent (attached to old stems for more than a season). |
Stems | usually 1–3, ascending to erect, scabrous to loosely puberulent. |
1–5+, erect, slender, glabrous. |
Leaves | basal and proximal usually withering by flowering, tapering to long-winged petioles, blades oblanceolate, 30–100 × 7–20(–30) mm, margins serrate or crenate, mid usually largest, apices acute to obtuse, acuminate, faces scabrous; mid and distal cauline subsessile (1 mm) or sessile, blades (sometimes ± shiny) elliptic to oblanceolate, 10–50 × 5–15(–25) mm, greatly reduced distally, grading into bracts, firm, bases convex-cuneate to rounded, margins finely serrate, often 3-nerved, nerves usually distinct abaxially, faces distinctly scabrous. |
basal tapering to long, winged petioles, blades ovate, to 7–22 mm wide, margins entire, apices acute, faces glabrous; proximal cauline tapering to winged petiole-like bases partially clasping stems, blades sometimes 3-nerved (2 prominent lateral nerves arising proximally and running alongside midnerve for some distance before diverging), linear oblanceolate, 158–177 × 7–22 mm, margins entire, ciliate, apices acute to obtuse, glabrous; mid to distal sessile, blades linear lanceolate, mid often 3-nerved (obscurely sometimes), distal 1-nerved, 47–100 × 4–10 mm, reduced distally. |
Peduncles | 0.5–2 mm; bracteoles 1–5, linear-lanceolate to ovate, minute, grading into phyllaries distally. |
5–6.2 mm, sparsely to moderately strigillose; bracteoles 0–2, linear. |
Involucres | narrowly campanulate, 3–5 mm. |
campanulate, (5.5–)6–8(–9) mm. |
Ray florets | 4–7; laminae 2–3.5 × 0.2–0.7 mm. |
6–12 (pale to bright yellow); laminae 7.4–7.9 × 0.5–0.6 mm. |
Disc florets | 4–6; corollas 3 mm, lobes 1 mm. |
8–15; corollas 4–4.3 mm, lobes 0.9–1.5 mm. |
Phyllaries | in 3–4 series, unequal, oblong, midnerves swollen distally, obtuse or acute to slightly acuminate. |
in 3–4 series, unequal, linear to oblanceolate, margins ciliate, weakly striate, obtuse to rounded. |
Heads | 20–260, in paniculiform arrays, narrowly to broadly secund, pyramidal, branches recurved, secund. |
(2–)5–50(–100+), not secund, in corymbiform arrays, branches and peduncles glabrous or moderately strigillose. |
Cypselae | 1.5–2.5 mm, sparsely to moderately short-strigose; pappi 3 mm. |
obconic, 1.4–1.8 mm, glabrous, ribs sometimes dark; pappi of bristles 4–5.5 mm, clavate. |
2n | = 18, 36. |
= 54. |
Solidago radula |
Solidago houghtonii |
|
Phenology | Flowering Aug–Oct. | Flowering Aug–Sep. |
Habitat | Open rocky places, dry woods, especially calcareous soils | Damp, interdunal hollows, limestone pavements (alvars) |
Elevation | 0–600 m (0–2000 ft) | 100–400 m (300–1300 ft) |
Distribution |
AR; GA; IL; KS; KY; LA; MO; NC; OK; SC; TX
|
MI; NY; ON |
Discussion | Solidago radula is disjunct in Georgia, North Carolina, and South Carolina. J. R. Beaudry (1969) reported a diploid from Smithville, Dekalb County, Tennessee; that has not been confirmed. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The large involucres and the presence of the unique 3-nerved proximal and mid cauline leaves distinguish Solidago houghtonii from other taxa in sect. Ptarmicoidei. The 3-nerved venation pattern may be morphologic evidence of a hybrid origin for S. houghtoni between a parallel-nerved member of sect. Ptarmicoidei and a 3-nerved member of subsect. Triplinerviae. Molecular data indicate that S. gigantea may have been one of the parental species (P. Laureto, pers. comm.). Reports of Solidago houghtonii from New York have been rejected as being based on misidentified specimens (J. K. Morton 1979). Some collections have the large involucres typical of S. houghtonii; molecular data indicate that the species is present in Genessee County, New York (P. Laureto, pers. comm.). Solidago houghtonii can be difficult to distinguish from S. ohioensis × S. ptarmicoides hybrids because both have hairy peduncles in arrays. The hybrids have pale yellow rays even when fresh; post-flowering rays of S. houghtonii can be pale yellow. J. K. Morton (1979) hypothesized an allopolyploid origin for the hexaploid S. houghtonii via an unreduced gamete of S. ohioensis × S. ptarmicoides backcrossed to S. ohioensis with subsequent chromosome doubling. Additional work is needed to confirm this hypothesis. An aberrant collection (M. R. Penskar 1067, MICH) had about 400 heads (mostly in bud) due to axillary branches developing along nearly the entire stem; stems usually have fewer than 50 heads. Some plants from Michigan are more robust than those found in Ontario. Solidago houghtonii is listed by the U.S. Fish and Wildlife Service as threatened in Michigan. It is in the Center for Plant Conservation’s National Collection of Endangered Plants. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 20, p. 162. | FNA vol. 20, p. 165. |
Parent taxa | Asteraceae > tribe Astereae > Solidago > sect. Solidago > subsect. Nemorales | Asteraceae > tribe Astereae > Solidago > sect. Ptarmicoidei |
Sibling taxa | ||
Synonyms | Aster decemflora, S. decemflora, S. laeta, S. pendula, S. radula var. laeta, S. radula var. rotundifolia, S. radula var. stenolepis, S. rotundifolia, S. scaberrima | Aster houghtonii, Oligoneuron houghtonii |
Name authority | Nuttall: J. Acad. Nat. Sci. Philadelphia 7: 102. (1834) | Torrey & A. Gray: in A. Gray, Manual, 211. (1848) |
Web links |