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downy goldenrod, downy ragged goldenrod

elm-leaf goldenrod

Habit Plants 40–150 cm; caudices stout, sometimes with long slender rhizomes. Plants 40–120 cm; caudices branching, woody.
Stems

1–20(–50+), stout, finely puberulent or scabrous-puberulent at least distally.

usually 1, erect, glabrous, sparsely hairy in arrays.

Leaves

basal absent at flowering;

cauline sessile to short-petiolate;

blades usually lanceolate-elliptic or ovate, sometimes linear-lanceolate, 30–150 × 5–30 mm, thick and firm, margins entire or few toothed, somewhat to much reduced distally, abaxial faces sometimes resinous and shiny, glabrous or strigillose (hairs mostly 0.1–1.4 mm), adaxial glabrous or scabrous.

basal when present similar to proximal cauline;

proximal cauline often withering by flowering, tapering (sometimes rather abruptly) to short, winged petioles, blades ovate-lanceolate, 60–100(–150) × 30–40(–50) mm, thin, margins coarsely serrate, apices acute, abaxial faces hirsute on main nerves, adaxial sparsely hirsute to somewhat scabrous;

mid to distal cauline subsessile to sessile, blades lanceolate, 20–50 × 5–20 mm, gradually reduced distally, margins entire.

Peduncles

mostly 2–15 mm, bracteolate, sparsely to densely short hispid-strigose.

1.7–2 mm, sparsely to moderately short hispido-strigose, bracteoles 2–7, ovate, grading into phyllaries.

Involucres

campanulate, 4.5–7.5 mm.

3–4 mm.

Ray florets

(5–)7–9;

laminae 3–7 × 1–2 mm.

3–6;

laminae 1–2 × 0.5–1 mm.

Disc florets

(8–)10–16;

corollas 4–5 mm, lobes ca. 1–2 mm.

4–7;

corollas 2.7–3 mm, lobes 0.5–1.1 mm.

Phyllaries

in 3–4 series, unequal, linear-lanceolate, acute to attenuate, ± squarrose-tipped, glabrous or moderately strigose, sparsely to moderately stipitate-glandular, sometimes viscid.

(16–18) in 2–3 series, unequal;

outer ovate, acute, inner linear-lanceolate, obtuse to acute.

Heads

10–190+ in paniculiform (rarely racemiform) arrays, usually elongate, usually leafy-bracteate, bracts similar to distal leaves but reduced;

branches stiffly ascending, not secund, sometimes elongate.

20–150, secund, in open paniculiform arrays, proximal branches elongate and widely divergent, sometimes pyramidal-secund with proximal branches short and recurved-secund.

Cypselae

3–4 mm, glabrous or glabrate;

pappi ca. 4 mm.

1–1.6 mm, finely hairy;

pappi ± 2.5 mm.

2n

= 18, 36, 54.

Solidago petiolaris

Solidago ulmifolia

Phenology Late Aug–Oct(–Nov).
Habitat Woods and open places, especially sandy soils
Elevation 0–1400[–2300] m (0–4600[–7500] ft)
Distribution
from FNA
AL; AR; FL; GA; IL; KS; LA; MO; NC; NE; NM; OK; SC; TX; Mexico (Coahuila)
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NE; NH; NJ; NY; OH; OK; PA; SC; TN; TX; VA; WI; WV; NS; ON
[WildflowerSearch map]
[BONAP county map]
Discussion

Solidago petiolaris is reported from Colorado but that report has not been confirmed. G. L. Nesom (1990j) discussed variation in the species and mapped the distribution. It is variable in leaf and phyllary shape and indument. Although several varieties have often been recognized in floras, the characters used to distinguish them form continua of variation that do not break into distinct groupings. Several general trends are worth noting. Plants in the Ozarks often have very resinous leaves; that does not appear to correlate with phyllary indument traits. Phyllary pubescence varies in a continuous fashion as well, with the numbers and distribution of hairs not breaking into discontinuous ranges. Diploids are known from throughout the range; one report of a tetraploid comes from North Carolina. Two reports of tetraploids from Florida were based on misidentified specimens. Plants with compact short arrays approach S. wrightii in this trait.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Solidago helleri is possibly a hybrid of S. ulmifolia with S. delicatula.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems glabrous or nearly so proximal to arrays; throughout range
var. ulmifolia
1. Stems distinctly spreading-hirsute; Arkansas, Mississippi, Missouri
var. palmeri
Source FNA vol. 20, p. 118. FNA vol. 20, p. 144.
Parent taxa Asteraceae > tribe Astereae > Solidago > sect. Solidago > subsect. Thyrsiflorae Asteraceae > tribe Astereae > Solidago > sect. Solidago > subsect. Venosae > ser. Venosae
Sibling taxa
S. albopilosa, S. altiplanities, S. altissima, S. arenicola, S. arguta, S. auriculata, S. bicolor, S. brachyphylla, S. buckleyi, S. caesia, S. canadensis, S. confinis, S. curtisii, S. delicatula, S. drummondii, S. elongata, S. erecta, S. faucibus, S. fistulosa, S. flexicaulis, S. gattingeri, S. gigantea, S. glomerata, S. guiradonis, S. hispida, S. houghtonii, S. juliae, S. juncea, S. kralii, S. lancifolia, S. latissimifolia, S. leavenworthii, S. leiocarpa, S. lepida, S. ludoviciana, S. macrophylla, S. missouriensis, S. mollis, S. multiradiata, S. nana, S. nemoralis, S. nitida, S. odora, S. ohioensis, S. ouachitensis, S. patula, S. pinetorum, S. plumosa, S. ptarmicoides, S. puberula, S. pulchra, S. radula, S. riddellii, S. rigida, S. roanensis, S. rugosa, S. rupestris, S. sciaphila, S. sempervirens, S. shortii, S. simplex, S. spathulata, S. speciosa, S. spectabilis, S. sphacelata, S. spithamaea, S. squarrosa, S. stricta, S. tarda, S. tortifolia, S. uliginosa, S. ulmifolia, S. velutina, S. verna, S. villosicarpa, S. wrightii
S. albopilosa, S. altiplanities, S. altissima, S. arenicola, S. arguta, S. auriculata, S. bicolor, S. brachyphylla, S. buckleyi, S. caesia, S. canadensis, S. confinis, S. curtisii, S. delicatula, S. drummondii, S. elongata, S. erecta, S. faucibus, S. fistulosa, S. flexicaulis, S. gattingeri, S. gigantea, S. glomerata, S. guiradonis, S. hispida, S. houghtonii, S. juliae, S. juncea, S. kralii, S. lancifolia, S. latissimifolia, S. leavenworthii, S. leiocarpa, S. lepida, S. ludoviciana, S. macrophylla, S. missouriensis, S. mollis, S. multiradiata, S. nana, S. nemoralis, S. nitida, S. odora, S. ohioensis, S. ouachitensis, S. patula, S. petiolaris, S. pinetorum, S. plumosa, S. ptarmicoides, S. puberula, S. pulchra, S. radula, S. riddellii, S. rigida, S. roanensis, S. rugosa, S. rupestris, S. sciaphila, S. sempervirens, S. shortii, S. simplex, S. spathulata, S. speciosa, S. spectabilis, S. sphacelata, S. spithamaea, S. squarrosa, S. stricta, S. tarda, S. tortifolia, S. uliginosa, S. velutina, S. verna, S. villosicarpa, S. wrightii
Subordinate taxa
S. ulmifolia var. palmeri, S. ulmifolia var. ulmifolia
Synonyms Aster lindheimeranus, Aster petiolaris, S. angusta, S. harperi, S. lindheimeriana, S. milleriana, S. petiolaris var. angusta, S. petiolaris var. squarrulosa, S. petiolaris var. wardii, S. squarrulosa, S. wardii Aster ulmifolius
Name authority Aiton: Hort. Kew. 3: 216. (1789) Muhlenberg ex Willdenow: Sp. Pl. 3: 2060. (1803)
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