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downy goldenrod, downy ragged goldenrod

Canada goldenrod, Rocky Mountain goldenrod, verge d'or élégante, western Canada goldenrod, western goldenrod

Habit Plants 40–150 cm; caudices stout, sometimes with long slender rhizomes. Plants 25–150 cm; rhizomes short to long creeping, forming few- to many-stemmed clones.
Stems

1–20(–50+), stout, finely puberulent or scabrous-puberulent at least distally.

1–25 (–50+), erect, proximally glabrous or sparsely to moderately canescent-villous, distally densely so.

Leaves

basal absent at flowering;

cauline sessile to short-petiolate;

blades usually lanceolate-elliptic or ovate, sometimes linear-lanceolate, 30–150 × 5–30 mm, thick and firm, margins entire or few toothed, somewhat to much reduced distally, abaxial faces sometimes resinous and shiny, glabrous or strigillose (hairs mostly 0.1–1.4 mm), adaxial glabrous or scabrous.

basal 0;

proximal cauline sessile, blades narrowly to broadly oblanceolate, 100–150 × 15–23 mm, tapering to bases, margins subentire to coarsely sharply serrate, 3-nerved, sometimes obscurely so, apices acute to acuminate, abaxial faces glabrate to sparsely short villous, adaxial glabrous;

mid to distal cauline sessile, blades lanceolate to narrowly elliptic, (50–)90–120(–150) × (7–) 11–23(–35) mm, largest near mid stem, then somewhat reduced distally in arrays, margins sharply and coarsely to finely serrate or entire, usually becoming less serrate near arrays, faces glabrous or sparsely strigoso-villous, more so along abaxial nerves, distalmost (below arrays) sometimes sparsely to moderately minutely stipitate-glandular.

Peduncles

mostly 2–15 mm, bracteolate, sparsely to densely short hispid-strigose.

1–4 mm, villoso-hirtellous, sometimes minutely stipitate-glandular;

bracteoles 0–2, linear-lanceolate, sometimes minutely stipitate-glandular.

Involucres

campanulate, 4.5–7.5 mm.

campanulate, (2–)2.3–3.8(–5) mm.

Ray florets

(5–)7–9;

laminae 3–7 × 1–2 mm.

(7–)10–16(–22);

laminae (0.5–)0.9–1.6(–2.2) × ca. 0.2–0.5 mm.

Disc florets

(8–)10–16;

corollas 4–5 mm, lobes ca. 1–2 mm.

(2–)5–9(–13);

corollas ca. (2–) 2.5–3.8(–5) mm, lobes (0.4–)0.6–1(–1.4) mm.

Phyllaries

in 3–4 series, unequal, linear-lanceolate, acute to attenuate, ± squarrose-tipped, glabrous or moderately strigose, sparsely to moderately stipitate-glandular, sometimes viscid.

in 3–4 series, deltate-lanceolate, unequal (outer 1/4–1/2 length of inner), margins sparsely villoso-ciliate, sometimes minutely stipitate-glandular apically or basally, apices acute to attenuate, glabrous or sparsely to moderately, minutely stipitate-glandular apically, sometimes over much of faces.

Heads

10–190+ in paniculiform (rarely racemiform) arrays, usually elongate, usually leafy-bracteate, bracts similar to distal leaves but reduced;

branches stiffly ascending, not secund, sometimes elongate.

14–800+, in sometimes secund, broadly thyrsiform or secund pyramidal arrays, usually leafy proximally, branches ascending or sometimes arching-spreading, leaves often minutely stipitate-glandular.

Cypselae

3–4 mm, glabrous or glabrate;

pappi ca. 4 mm.

(narrowly obconic) 0.6–1.2 mm, strigillose;

pappi 2.5–3.3 mm.

2n

= 18, 36, 54.

Solidago petiolaris

Solidago lepida

Phenology Late Aug–Oct(–Nov).
Habitat Woods and open places, especially sandy soils
Elevation 0–1400[–2300] m (0–4600[–7500] ft)
Distribution
from FNA
AL; AR; FL; GA; IL; KS; LA; MO; NC; NE; NM; OK; SC; TX; Mexico (Coahuila)
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AZ; CA; CO; ID; MT; NM; NV; UT; WA; WY; AB; BC; MB; NB; NF; NT; ON; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Solidago petiolaris is reported from Colorado but that report has not been confirmed. G. L. Nesom (1990j) discussed variation in the species and mapped the distribution. It is variable in leaf and phyllary shape and indument. Although several varieties have often been recognized in floras, the characters used to distinguish them form continua of variation that do not break into distinct groupings. Several general trends are worth noting. Plants in the Ozarks often have very resinous leaves; that does not appear to correlate with phyllary indument traits. Phyllary pubescence varies in a continuous fashion as well, with the numbers and distribution of hairs not breaking into discontinuous ranges. Diploids are known from throughout the range; one report of a tetraploid comes from North Carolina. Two reports of tetraploids from Florida were based on misidentified specimens. Plants with compact short arrays approach S. wrightii in this trait.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Subspecies 2 (2 in the flora).

Solidago lepida is the only member of subsect. Triplinerviae that is nearly always minutely, sometimes sparsely, stipitate-glandular in the arrays. These glands can be very small and visible only with 30–70× magnification. The enlarged head of the stalked gland may be little bigger than a pollen grain. The species occurs in the west from the Aleutian Islands and central Alaska south just into northern California, and in the Rocky Mountains to Arizona and New Mexico. Its range extends across Canada to Newfoundland, the Gaspé (Quebec), and northern New Brunswick. It also appears to be introduced at scattered locations across the Canadian prairies.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Distal cauline leaves usually with 6–8 teeth per side; phyllaries very sparsely, minutely stipitate-glandular along proximal margins; e Canada (Solidago lepida subsp. fallax)
subsp. fallax
1. Distal cauline leaves usually with 0–6 teeth per side (plants of w mountains in United States sometimes more serrate); phyllaries, peduncles, peduncle bracts and/or distalmost cauline leaves sparsely to moderately, minutely stipitate-glandular; mountains, Alaska to Arizona and New Mexico, and across Canada, scattered eastward to the Gaspé, Quebec (S. lepida subsp. lepida)
→ 2
2. Heads in thyrsiform arrays, proximal branches ascending; much of range
var. lepida
2. Heads in pyramidal paniculiform arrays, proximal branches arching to recurved; Rocky Mountains and Cascade Mountains, Alberta and British Columbia to California, Arizona, New Mexico
var. salebrosa
Source FNA vol. 20, p. 118. FNA vol. 20, p. 155.
Parent taxa Asteraceae > tribe Astereae > Solidago > sect. Solidago > subsect. Thyrsiflorae Asteraceae > tribe Astereae > Solidago > sect. Solidago > subsect. Triplinerviae
Sibling taxa
S. albopilosa, S. altiplanities, S. altissima, S. arenicola, S. arguta, S. auriculata, S. bicolor, S. brachyphylla, S. buckleyi, S. caesia, S. canadensis, S. confinis, S. curtisii, S. delicatula, S. drummondii, S. elongata, S. erecta, S. faucibus, S. fistulosa, S. flexicaulis, S. gattingeri, S. gigantea, S. glomerata, S. guiradonis, S. hispida, S. houghtonii, S. juliae, S. juncea, S. kralii, S. lancifolia, S. latissimifolia, S. leavenworthii, S. leiocarpa, S. lepida, S. ludoviciana, S. macrophylla, S. missouriensis, S. mollis, S. multiradiata, S. nana, S. nemoralis, S. nitida, S. odora, S. ohioensis, S. ouachitensis, S. patula, S. pinetorum, S. plumosa, S. ptarmicoides, S. puberula, S. pulchra, S. radula, S. riddellii, S. rigida, S. roanensis, S. rugosa, S. rupestris, S. sciaphila, S. sempervirens, S. shortii, S. simplex, S. spathulata, S. speciosa, S. spectabilis, S. sphacelata, S. spithamaea, S. squarrosa, S. stricta, S. tarda, S. tortifolia, S. uliginosa, S. ulmifolia, S. velutina, S. verna, S. villosicarpa, S. wrightii
S. albopilosa, S. altiplanities, S. altissima, S. arenicola, S. arguta, S. auriculata, S. bicolor, S. brachyphylla, S. buckleyi, S. caesia, S. canadensis, S. confinis, S. curtisii, S. delicatula, S. drummondii, S. elongata, S. erecta, S. faucibus, S. fistulosa, S. flexicaulis, S. gattingeri, S. gigantea, S. glomerata, S. guiradonis, S. hispida, S. houghtonii, S. juliae, S. juncea, S. kralii, S. lancifolia, S. latissimifolia, S. leavenworthii, S. leiocarpa, S. ludoviciana, S. macrophylla, S. missouriensis, S. mollis, S. multiradiata, S. nana, S. nemoralis, S. nitida, S. odora, S. ohioensis, S. ouachitensis, S. patula, S. petiolaris, S. pinetorum, S. plumosa, S. ptarmicoides, S. puberula, S. pulchra, S. radula, S. riddellii, S. rigida, S. roanensis, S. rugosa, S. rupestris, S. sciaphila, S. sempervirens, S. shortii, S. simplex, S. spathulata, S. speciosa, S. spectabilis, S. sphacelata, S. spithamaea, S. squarrosa, S. stricta, S. tarda, S. tortifolia, S. uliginosa, S. ulmifolia, S. velutina, S. verna, S. villosicarpa, S. wrightii
Subordinate taxa
S. lepida subsp. fallax, S. lepida var. lepida, S. lepida var. salebrosa
Synonyms Aster lindheimeranus, Aster petiolaris, S. angusta, S. harperi, S. lindheimeriana, S. milleriana, S. petiolaris var. angusta, S. petiolaris var. squarrulosa, S. petiolaris var. wardii, S. squarrulosa, S. wardii Aster lepidus, S. canadensis var. lepida
Name authority Aiton: Hort. Kew. 3: 216. (1789) de Candolle: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 5: 339. (1836)
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