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broad-leaf goldenrod, verge d'or à tige zigzaguante, zig-zag goldenrod

downy goldenrod, downy ragged goldenrod

Habit Plants (15–)25–75(–90) cm; caudices woody, rhizomes short. Plants 40–150 cm; caudices stout, sometimes with long slender rhizomes.
Stems

1–5+, flexuous, sparsely to moderately hairy in arrays.

1–20(–50+), stout, finely puberulent or scabrous-puberulent at least distally.

Leaves

rosettes present at flowering;

basal and proximal cauline tapering to winged petioles, blades ovate, (70–)100–140(–180) × (27–)40–60(–80) mm (petioles 1/4–1/2 total leaf length), margins serrate-serrulate [teeth (14–)21–33(–45)], abaxial faces glabrous or moderately hairy, more densely so on nerves, adaxial glabrous or sparsely hairy;

mid to distal cauline sessile, blades narrowly ovate to lanceolate, 38–90(–150) × 10–30(–55) mm, reduced distally, becoming lanceolate in arrays, bases tapering, margins entire to serrate, apices acuminate to cuspidate, faces glabrous or sparsely hairy, abaxial glabrous or moderately hairy, more densely so along nerves.

basal absent at flowering;

cauline sessile to short-petiolate;

blades usually lanceolate-elliptic or ovate, sometimes linear-lanceolate, 30–150 × 5–30 mm, thick and firm, margins entire or few toothed, somewhat to much reduced distally, abaxial faces sometimes resinous and shiny, glabrous or strigillose (hairs mostly 0.1–1.4 mm), adaxial glabrous or scabrous.

Peduncles

0.5–5 mm, moderately to densely strigose;

bracteoles 1–3, lanceolate, usually near base of involucres and grading into phyllaries.

mostly 2–15 mm, bracteolate, sparsely to densely short hispid-strigose.

Involucres

campanulate, 4.5–7(–8) mm.

campanulate, 4.5–7.5 mm.

Ray florets

1–5;

laminae (2–)2.5–4(–5) × 0.7–2 mm.

(5–)7–9;

laminae 3–7 × 1–2 mm.

Disc florets

4–8(–11);

corollas 2–3(–4) mm, lobes 1–1.6(–2) mm.

(8–)10–16;

corollas 4–5 mm, lobes ca. 1–2 mm.

Phyllaries

in ca. 3 series, strongly unequal, outermost 1–2 mm, innermost 3.7–4.7(–5.7) mm, linear-oblong, 1-nerved, apices obtuse to acute.

in 3–4 series, unequal, linear-lanceolate, acute to attenuate, ± squarrose-tipped, glabrous or moderately strigose, sparsely to moderately stipitate-glandular, sometimes viscid.

Heads

25–250, in short axillary and terminal racemiform clusters, lateral panicles (2–)7–31(–56) cm.

10–190+ in paniculiform (rarely racemiform) arrays, usually elongate, usually leafy-bracteate, bracts similar to distal leaves but reduced;

branches stiffly ascending, not secund, sometimes elongate.

Cypselae

(obconic) 1–2(–3) mm, moderately to densely strigose;

pappi 3–4.5 mm.

3–4 mm, glabrous or glabrate;

pappi ca. 4 mm.

2n

= 18, 36.

= 18, 36, 54.

Solidago flexicaulis

Solidago petiolaris

Phenology Flowering Aug–Oct. Late Aug–Oct(–Nov).
Habitat Shaded woods and thickets Woods and open places, especially sandy soils
Elevation 0–1200+ m (0–3900+ ft) 0–1400[–2300] m (0–4600[–7500] ft)
Distribution
from FNA
AL; AR; CT; DE; GA; IA; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; NC; ND; NE; NH; NJ; NY; OH; PA; SD; TN; VA; VT; WI; WV; NB; NS; ON; PE; QC
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; FL; GA; IL; KS; LA; MO; NC; NE; NM; OK; SC; TX; Mexico (Coahuila)
[WildflowerSearch map]
[BONAP county map]
Discussion

The two cytotypes of Solidago flexicaulis show a strong geographic pattern. Diploids mostly occur east of the Appalachians except in the southwestern portion of the range, while tetraploids occur west of the mountains (J. G. Chmielewski and J. C. Semple 1985). The report of 2n = 90 for the species (Semple et al. 1993) was based on a specimen of the recently described Solidago faucibus. The significance of ploidy level on cypselae traits was analyzed in detail by Chmielewski et al. (1989).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Solidago petiolaris is reported from Colorado but that report has not been confirmed. G. L. Nesom (1990j) discussed variation in the species and mapped the distribution. It is variable in leaf and phyllary shape and indument. Although several varieties have often been recognized in floras, the characters used to distinguish them form continua of variation that do not break into distinct groupings. Several general trends are worth noting. Plants in the Ozarks often have very resinous leaves; that does not appear to correlate with phyllary indument traits. Phyllary pubescence varies in a continuous fashion as well, with the numbers and distribution of hairs not breaking into discontinuous ranges. Diploids are known from throughout the range; one report of a tetraploid comes from North Carolina. Two reports of tetraploids from Florida were based on misidentified specimens. Plants with compact short arrays approach S. wrightii in this trait.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 20, p. 128. FNA vol. 20, p. 118.
Parent taxa Asteraceae > tribe Astereae > Solidago > sect. Solidago > subsect. Glomeruliflorae Asteraceae > tribe Astereae > Solidago > sect. Solidago > subsect. Thyrsiflorae
Sibling taxa
S. albopilosa, S. altiplanities, S. altissima, S. arenicola, S. arguta, S. auriculata, S. bicolor, S. brachyphylla, S. buckleyi, S. caesia, S. canadensis, S. confinis, S. curtisii, S. delicatula, S. drummondii, S. elongata, S. erecta, S. faucibus, S. fistulosa, S. gattingeri, S. gigantea, S. glomerata, S. guiradonis, S. hispida, S. houghtonii, S. juliae, S. juncea, S. kralii, S. lancifolia, S. latissimifolia, S. leavenworthii, S. leiocarpa, S. lepida, S. ludoviciana, S. macrophylla, S. missouriensis, S. mollis, S. multiradiata, S. nana, S. nemoralis, S. nitida, S. odora, S. ohioensis, S. ouachitensis, S. patula, S. petiolaris, S. pinetorum, S. plumosa, S. ptarmicoides, S. puberula, S. pulchra, S. radula, S. riddellii, S. rigida, S. roanensis, S. rugosa, S. rupestris, S. sciaphila, S. sempervirens, S. shortii, S. simplex, S. spathulata, S. speciosa, S. spectabilis, S. sphacelata, S. spithamaea, S. squarrosa, S. stricta, S. tarda, S. tortifolia, S. uliginosa, S. ulmifolia, S. velutina, S. verna, S. villosicarpa, S. wrightii
S. albopilosa, S. altiplanities, S. altissima, S. arenicola, S. arguta, S. auriculata, S. bicolor, S. brachyphylla, S. buckleyi, S. caesia, S. canadensis, S. confinis, S. curtisii, S. delicatula, S. drummondii, S. elongata, S. erecta, S. faucibus, S. fistulosa, S. flexicaulis, S. gattingeri, S. gigantea, S. glomerata, S. guiradonis, S. hispida, S. houghtonii, S. juliae, S. juncea, S. kralii, S. lancifolia, S. latissimifolia, S. leavenworthii, S. leiocarpa, S. lepida, S. ludoviciana, S. macrophylla, S. missouriensis, S. mollis, S. multiradiata, S. nana, S. nemoralis, S. nitida, S. odora, S. ohioensis, S. ouachitensis, S. patula, S. pinetorum, S. plumosa, S. ptarmicoides, S. puberula, S. pulchra, S. radula, S. riddellii, S. rigida, S. roanensis, S. rugosa, S. rupestris, S. sciaphila, S. sempervirens, S. shortii, S. simplex, S. spathulata, S. speciosa, S. spectabilis, S. sphacelata, S. spithamaea, S. squarrosa, S. stricta, S. tarda, S. tortifolia, S. uliginosa, S. ulmifolia, S. velutina, S. verna, S. villosicarpa, S. wrightii
Synonyms Doria flexicaulis, S. flexicaulis var. ciliata, S. flexicaulis var. latifolia, S. latifolia, S. scrophulariifolia Aster lindheimeranus, Aster petiolaris, S. angusta, S. harperi, S. lindheimeriana, S. milleriana, S. petiolaris var. angusta, S. petiolaris var. squarrulosa, S. petiolaris var. wardii, S. squarrulosa, S. wardii
Name authority Linnaeus: Sp. Pl. 2: 879. (1753) Aiton: Hort. Kew. 3: 216. (1789)
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