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Buckley's goldenrod

alpine goldenrod, dune goldenrod, Mt. Albert goldenrod, Rand's goldenrod, spikelike goldenrod, sticky goldenrod

Habit Plants 60–120 cm; caudices thick, woody, roots thick. Plants 5–80 cm; caudices branching.
Stems

1–5+, sparsely to moderately short strigose or villous.

1–10+, ascending to erect, proximally glabrous, strigose in arrays.

Leaves

basal withering by flowering, petiolate, smaller to much smaller than cauline, blades oblanceolate, margins serrate;

cauline sessile, blades elliptic-lanceolate or -oblanceolate, mid 80–140 × 2.5–4 mm, distally reduced, usually membranous, bases tapering, attenuate, margins sharply toothed along much of length to nearly entire, ciliate, abaxial faces short-pilose along small and large nerves, adaxial short-pilose along larger nerves.

basal and proximal cauline petiolate, blades narrowly oblanceolate, (20–)50–100(–160) × (2–)5–16(–56) mm, margins serrate to crenate, apices acute to obtuse, glabrous;

mid and distal sessile, similar, blades lanceolate to linear, 12–45 × 2–19 mm, reduced distally, margins entire to sparsely serrate, sometimes resinous.

Peduncles

1–6 mm, moderately canescent;

bracteoles linear to lanceolate.

3.1–10.3 mm, glabrate to sparsely strigillose;

bracteoles few, linear.

Involucres

campanulate, 4.5–5.5 mm.

campanulate, 3–7 mm.

Ray florets

6–8;

laminae 3–4 × 1–1.5 mm wide.

7–16;

laminae 2–5 × 0.7–0.9 mm.

Disc florets

8–14;

corollas 4–5 mm, lobes ca. 1.5 mm.

6–31;

corollas 4–4.9 mm, lobes 0.6–1.3(–2) mm.

Phyllaries

in ca. 3 series, strongly unequal, erect to slightly squarrose-tipped, lanceolate, apices acute, glabrate, sparsely to moderately, finely stipitate-glandular.

(in 3–4 series) strongly unequal, often resinous;

outer ovate, acute, inner linear-oblong, obtuse.

Heads

5–160 (2–10+ more per short branch cluster), in narrowly elongate paniculiform arrays, branches usually 1–6 cm (much longer in damaged plants, sometimes 1–3 proximal branches much elongated in undamaged plants), ascending and bearing short terminal racemiform or paniculiform clusters.

3–150, not secund, in narrowly elongate, paniculiform arrays, broadly so in robust plants (12.5–19 × 2.5–3 cm wide), consisting of short axillary and terminal racemiform clusters, proximal branches elongate in larger plants, branches glabrate to strigillose.

Cypselae

(reddish brown) 2–3 mm, glabrous;

pappi 4–5 mm.

narrowly obconic, 1.9–3.2 mm, sometimes with dark ridges, strigillose;

pappi 1.9–5.2 mm (bristles sometimes clavate).

Solidago buckleyi

Solidago simplex

Phenology Flowering Sep.
Habitat Open oak woods, ridges and slopes, bluffs
Elevation 100–300 m (300–1000 ft)
Distribution
from FNA
AR; IL; IN; KY; MO
[WildflowerSearch map]
[BONAP county map]
from FNA
AK; AZ; CO; ID; IN; MA; MD; ME; MI; MN; MT; ND; NH; NM; NY; OR; PA; SD; UT; VT; WA; WI; WV; WY; AB; BC; MB; NB; NT; ON; QC; SK; YT; Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Solidago buckleyi is an uncommon species of mesic woods, most variable in the size and number of teeth on the large mid cauline leaves. Once seen, usually it is not easily confused with S. petiolaris. Reports from farther east are for plants of S. petiolaris.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 7 (7 in the flora).

The somewhat viscid-resinous heads of Solidago simplex are its most distinctive feature, separating it from similar sympatric species. G. S. Ringius (1985) did a detailed multivariate analysis of the S. spathulata/S. simplex complex (the latter under the name S. glutinosa). The cytogeography of the species complex was presented by Ringius and J. C. Semple (1987). Neither study included data on the next three species occurring in the southeastern United States.

The species is divided into two subspecies and seven varieties following G. S Ringius (1985) and J. C. Semple et al. (1999). Three varieties occur in the diploid transcontinental subsp. simplex: var. simplex, var. nana, and var. chlorolepis. Four varieties occur in the eastern North American tetraploid-hexaploid subsp. randii: var. monticola, var. gillmanii, var. ontarioensis, and var. racemosa. Except for var. simplex, varieties are restricted to different habitats in relatively limited ranges.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Basal and cauline leaves spatulate to orbiculate, apices obtuse to rounded; cypselae moderately to densely strigose (S. simplex subsp. simplex)
→ 2
1. Basal and cauline leaves spatulate to obovate, apices usually acute; cypselae sparsely to moderately strigose (S. simplex subsp. randii)
→ 5
2. Cauline leaves 5–16 (longest 60+ mm); w North America, disjunct along shores of upper Great Lakes and in s Quebec
var. simplex
2. Cauline leaves 2–7 (longest to 60 mm); alpine slopes, w North America, Mt. Albert, Gaspé, Quebec
→ 3
3. Shortest phyllaries 1.4–2 mm; disc corollas 3.9–4.2 mm; serpentine soils, Mt. Albert, Gaspé, Quebec
var. chlorolepis
3. Shortest phyllaries 2.1–3.1 mm; disc corollas 4.2–5.4 mm; alpine slopes, Rocky Mountains, Cascade Mountains, Vancouver Island.
→ 4
4. Basal leaves usually spatulate, rarely broadly elliptic, proximal blades gradually attenuated to winged petioles, margins weakly crenate; alpine slopes, Rocky Mountains
var. simplex
4. Basal leaves orbiculate to broadly elliptic, proximal blades abruptly attenuated to winged petioles, margins distinctly crenate; alpine slopes of Cascade Mountains, Vancouver Island
var. nana
5. Proximal cauline leaves spatulate to obovate, (5–)10–42 mm wide; leaf margins often sharply serrate
→ 6
5. Proximal cauline leaves lanceolate to narrowly spatulate, 2–10(–21.5) mm wide; leaf margins entire or dentate, rarely serrate
→ 7
6. Plants (10–)18–53(–83) cm, arrays tending to be compact, virgate; proximal cauline leaves (5–)10–22(–31) mm wide, margins crenate to serrate; barren, rocky, non-alpine uplands; se Quebec, New England, New York
var. monticola
6. Plants (20–)38–73(–84) cm, robust, arrays tending to be paniculiform; proximal cauline leaves (6–)7.5–24(–42) mm wide, margins often sharply serrate; sand dunes, Lake Michigan, n Lake Huron
var. gillmanii
7. Cauline leaves (3–)12–33(–58); peduncle bracteoles 2+; calcareous rocky riverbanks, e Quebec s to West Virginia and Maryland
var. racemosa
7. Cauline leaves (2–)4–13(–24); peduncle bracteoles 1–3(–4); rocky shores, Lake Superior, n Lake Huron
var. ontarioensis
Source FNA vol. 20, p. 118. FNA vol. 20, p. 113.
Parent taxa Asteraceae > tribe Astereae > Solidago > sect. Solidago > subsect. Thyrsiflorae Asteraceae > tribe Astereae > Solidago > sect. Solidago > subsect. Humiles
Sibling taxa
S. albopilosa, S. altiplanities, S. altissima, S. arenicola, S. arguta, S. auriculata, S. bicolor, S. brachyphylla, S. caesia, S. canadensis, S. confinis, S. curtisii, S. delicatula, S. drummondii, S. elongata, S. erecta, S. faucibus, S. fistulosa, S. flexicaulis, S. gattingeri, S. gigantea, S. glomerata, S. guiradonis, S. hispida, S. houghtonii, S. juliae, S. juncea, S. kralii, S. lancifolia, S. latissimifolia, S. leavenworthii, S. leiocarpa, S. lepida, S. ludoviciana, S. macrophylla, S. missouriensis, S. mollis, S. multiradiata, S. nana, S. nemoralis, S. nitida, S. odora, S. ohioensis, S. ouachitensis, S. patula, S. petiolaris, S. pinetorum, S. plumosa, S. ptarmicoides, S. puberula, S. pulchra, S. radula, S. riddellii, S. rigida, S. roanensis, S. rugosa, S. rupestris, S. sciaphila, S. sempervirens, S. shortii, S. simplex, S. spathulata, S. speciosa, S. spectabilis, S. sphacelata, S. spithamaea, S. squarrosa, S. stricta, S. tarda, S. tortifolia, S. uliginosa, S. ulmifolia, S. velutina, S. verna, S. villosicarpa, S. wrightii
S. albopilosa, S. altiplanities, S. altissima, S. arenicola, S. arguta, S. auriculata, S. bicolor, S. brachyphylla, S. buckleyi, S. caesia, S. canadensis, S. confinis, S. curtisii, S. delicatula, S. drummondii, S. elongata, S. erecta, S. faucibus, S. fistulosa, S. flexicaulis, S. gattingeri, S. gigantea, S. glomerata, S. guiradonis, S. hispida, S. houghtonii, S. juliae, S. juncea, S. kralii, S. lancifolia, S. latissimifolia, S. leavenworthii, S. leiocarpa, S. lepida, S. ludoviciana, S. macrophylla, S. missouriensis, S. mollis, S. multiradiata, S. nana, S. nemoralis, S. nitida, S. odora, S. ohioensis, S. ouachitensis, S. patula, S. petiolaris, S. pinetorum, S. plumosa, S. ptarmicoides, S. puberula, S. pulchra, S. radula, S. riddellii, S. rigida, S. roanensis, S. rugosa, S. rupestris, S. sciaphila, S. sempervirens, S. shortii, S. spathulata, S. speciosa, S. spectabilis, S. sphacelata, S. spithamaea, S. squarrosa, S. stricta, S. tarda, S. tortifolia, S. uliginosa, S. ulmifolia, S. velutina, S. verna, S. villosicarpa, S. wrightii
Subordinate taxa
S. simplex var. chlorolepis, S. simplex var. gillmanii, S. simplex var. monticola, S. simplex var. nana, S. simplex var. ontarioensis, S. simplex var. racemosa, S. simplex var. simplex
Synonyms Aster buckleyi
Name authority Torrey & A. Gray: Fl. N. Amer. 2: 198. (1842) Kunth: in A. von Humboldt et al., Nov. Gen. Sp. 4(fol.): 81. (1818)
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