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tropical soda apple

twinleaf nightshade, twoleaf nightshade

Habit Shrubs, erect, sparsely armed, 0.5–2 m, prickles white or yellowish, straight or recurved, 1–25 mm, densely pubescent, hairs unbranched, short-glandular and longer-eglandular, with sessile, stellate hairs on abaxial leaf surface, these 4(–5)-rayed, central ray shorter than lateral rays. Shrubs, erect, unarmed, 1–2 m, glabrous or occasionally minutely puberulent, hairs white, unbranched, eglandular.
Leaves

petiolate;

petiole 3–6 cm;

blade simple, ovate to suborbiculate, 7–10(–20) × 6–8(–15) cm, margins coarsely lobed with 3–5 lobes per side, lobe margins entire to coarsely toothed, base truncate to cordate.

petiolate;

petiole 0.2–0.5 cm;

blade simple, elliptic, 0.9–6.8 × 0.6–2.2 cm, margins entire, base acute to attenuate or decurrent.

Inflorescences

extra-axillary, sessile or nearly so, unbranched, 3–5-flowered.

leaf-opposed, unbranched, 5–20-flowered, 0.3–1.2 cm.

Pedicels

0.7–1.1 cm in flower, 1–2 cm in fruit.

ca. 0.5 cm in flower, ca. 1.2 cm and erect in fruit.

Flowers

radially symmetric;

calyx somewhat accrescent, unarmed or sparsely prickly, 3–4 mm, densely pubescent, lobes triangular;

corolla greenish or whitish, stellate, 1.5–2.5 cm diam., without interpetalar tissue;

stamens equal;

anthers narrow and tapered, 5.5–7(–10) mm, dehiscent by terminal pores;

ovary densely pubescent, hairs glandular and eglandular.

radially symmetric;

calyx somewhat accrescent, unarmed, 1.5–2 mm, glabrous, lobes deltoid;

corolla white, often tinged with lavender, stellate, 0.7–1 cm diam., without interpetalar tissue;

stamens equal;

anthers oblong, 1.5–2 mm, dehiscent by terminal pores that open into longitudinal slits;

ovary glabrous.

Berries

light green mottled with dark green when young, yellow when ripe, globose, (1.5–)2–3 cm diam., glabrous, without sclerotic granules.

yellow to orange, globose, 0.7–1.2 cm diam., glabrous, without sclerotic granules.

Seeds

reddish brown, flattened, 2–3 × 2–2.5 mm, minutely pitted.

pale yellow or tan, flattened, ca. 3 × 2.5 mm, minutely pitted.

2n

= 24.

= 24.

Solanum viarum

Solanum diphyllum

Phenology Flowering May–frost (year-round in Fla.). Flowering year-round.
Habitat Pastures, roadsides, disturbed areas. Dry lowland areas, ham­mocks, disturbed sites.
Elevation 0–1000 m. (0–3300 ft.) 0–300 m. (0–1000 ft.)
Distribution
from FNA
AL; FL; GA; LA; MS; NC; PA; SC; TN; TX; South America (Argentina, Brazil, Paraguay, Uruguay); intro­duced also in Asia (India); Africa [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from FNA
FL; Mexico; West Indies; Central America (Belize, Costa Rica, El Salvador, Guatemala, Honduras, Nicaragua); South America (Brazil) [Introduced in North America; introduced also in Europe (s France, Italy), Asia, Pacific Islands (Java, Philippines)]
[WildflowerSearch map]
[BONAP county map]
Discussion

In the United States, Solanum viarum was first collected in Florida in 1988 and has subsequently become an aggressive and invasive species in the Southeast. It is on the Federal Noxious Weeds List and is classified as a noxious weed or plant pest in many states. Cattle and other animals eat the fruits and spread the seeds through their feces, and the seeds are coated with a sticky substance that makes them adhere to farm equipment when the plants are mowed. It can form large patches that are difficult to eradicate due to their extensive root systems and sharp prickles. It is a major agricultural pest and a threat to native ecosystems.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Solanum diphyllum is often cultivated for its brightly colored fruits and can escape from cultivation in tropical and subtropical areas. It occurs sporadically and does not appear to be common, but in other areas where it has escaped it has become naturalized (Asia; S. Knapp 2002b).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 14. FNA vol. 14.
Parent taxa Solanaceae > Solanum Solanaceae > Solanum
Sibling taxa
S. americanum, S. aviculare, S. bahamense, S. campechiense, S. capsicoides, S. carolinense, S. chenopodioides, S. citrullifolium, S. cordicitum, S. davisense, S. deflexum, S. dimidiatum, S. diphyllum, S. donianum, S. douglasii, S. dulcamara, S. elaeagnifolium, S. emulans, S. erianthum, S. furcatum, S. hindsianum, S. interius, S. jamaicense, S. jamesii, S. laciniatum, S. lanceolatum, S. lumholtzianum, S. lycopersicum, S. marginatum, S. mauritianum, S. nigrescens, S. nigrum, S. nitidibaccatum, S. novomexicanum, S. perplexum, S. pseudocapsicum, S. pseudogracile, S. pumilum, S. rostratum, S. sarrachoides, S. seaforthianum, S. setigeroides, S. sisymbriifolium, S. stoloniferum, S. tampicense, S. tenuipes, S. torvum, S. triflorum, S. triquetrum, S. umbelliferum, S. wallacei
S. americanum, S. aviculare, S. bahamense, S. campechiense, S. capsicoides, S. carolinense, S. chenopodioides, S. citrullifolium, S. cordicitum, S. davisense, S. deflexum, S. dimidiatum, S. donianum, S. douglasii, S. dulcamara, S. elaeagnifolium, S. emulans, S. erianthum, S. furcatum, S. hindsianum, S. interius, S. jamaicense, S. jamesii, S. laciniatum, S. lanceolatum, S. lumholtzianum, S. lycopersicum, S. marginatum, S. mauritianum, S. nigrescens, S. nigrum, S. nitidibaccatum, S. novomexicanum, S. perplexum, S. pseudocapsicum, S. pseudogracile, S. pumilum, S. rostratum, S. sarrachoides, S. seaforthianum, S. setigeroides, S. sisymbriifolium, S. stoloniferum, S. tampicense, S. tenuipes, S. torvum, S. triflorum, S. triquetrum, S. umbelliferum, S. viarum, S. wallacei
Name authority Dunal in A. P. de Candolle and A. L. P. P. de Candolle: Prodr. 13(1): 240. (1852) Linnaeus: Sp. Pl. 1: 184. (1753)
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