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tropical soda apple

apple of sodom, bull nettle, Carolina horse-nettle, Carolina nightshade, devil's tomato, horse nettle

Habit Shrubs, erect, sparsely armed, 0.5–2 m, prickles white or yellowish, straight or recurved, 1–25 mm, densely pubescent, hairs unbranched, short-glandular and longer-eglandular, with sessile, stellate hairs on abaxial leaf surface, these 4(–5)-rayed, central ray shorter than lateral rays. Herbs or shrubs, perennial, erect, sparsely to moderately armed, to 1.2 m, prickles cream to yellowish, straight, to 6 mm, sparsely to densely pubescent, hairs white to cream, sessile to short-stalked, stellate, 4(–8)-rayed, central ray 1–3(–5)-celled and to 3 mm, longer than lateral rays.
Leaves

petiolate;

petiole 3–6 cm;

blade simple, ovate to suborbiculate, 7–10(–20) × 6–8(–15) cm, margins coarsely lobed with 3–5 lobes per side, lobe margins entire to coarsely toothed, base truncate to cordate.

petiolate;

petiole 0.4–4 cm;

blade simple, ovate, lanceolate, or elliptic, 2–15 × 2–10 cm, margins subentire, sinuate, or lobed with 1–4 lobes per side, sometimes very deeply lobed almost to midrib, lobe margins entire to coarsely lobed, base cuneate.

Inflorescences

extra-axillary, sessile or nearly so, unbranched, 3–5-flowered.

extra-axillary, unbranched or rarely forked, 2–12-flowered, 2–9 cm.

Pedicels

0.7–1.1 cm in flower, 1–2 cm in fruit.

0.5–1 cm in flower, 1.2–1.8 cm and curved downward in fruit.

Flowers

radially symmetric;

calyx somewhat accrescent, unarmed or sparsely prickly, 3–4 mm, densely pubescent, lobes triangular;

corolla greenish or whitish, stellate, 1.5–2.5 cm diam., without interpetalar tissue;

stamens equal;

anthers narrow and tapered, 5.5–7(–10) mm, dehiscent by terminal pores;

ovary densely pubescent, hairs glandular and eglandular.

radially symmetric;

calyx somewhat accrescent, unarmed or with sparse prickles, 5–8 mm, moderately stellate-pubescent, lobes lanceolate to elliptic-acuminate;

corolla white to pale blue or violet, stellate to stellate-pentagonal or rotate-stellate, 2–3 cm diam., with interpetalar tissue at margins and bases of lobes;

stamens equal;

anthers narrow and tapered, 4.5–6.5 mm, dehiscent by terminal pores;

ovary glabrous or sparsely to moderately glandular-puberulent, rarely moderately pubescent, hairs white, stellate or simple.

Berries

light green mottled with dark green when young, yellow when ripe, globose, (1.5–)2–3 cm diam., glabrous, without sclerotic granules.

light green with darker green mottling or pale greenish white, turning bright yellow, subglobose to depressed-globose, 1–2 × 1–1.8 cm, glabrous, without sclerotic granules.

Seeds

reddish brown, flattened, 2–3 × 2–2.5 mm, minutely pitted.

yellow, flattened, 1.7–2.4 × 1.6–1.8 mm, minutely pitted.

2n

= 24.

= 24.

Solanum viarum

Solanum carolinense

Phenology Flowering May–frost (year-round in Fla.).
Habitat Pastures, roadsides, disturbed areas.
Elevation 0–1000 m. (0–3300 ft.)
Distribution
from FNA
AL; FL; GA; LA; MS; NC; PA; SC; TN; TX; South America (Argentina, Brazil, Paraguay, Uruguay); intro­duced also in Asia (India); Africa [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
from USDA
North America [Introduced in Europe, Asia, Pacific Islands (New Zealand), Australia]
[WildflowerSearch map]
[BONAP county map]
Discussion

In the United States, Solanum viarum was first collected in Florida in 1988 and has subsequently become an aggressive and invasive species in the Southeast. It is on the Federal Noxious Weeds List and is classified as a noxious weed or plant pest in many states. Cattle and other animals eat the fruits and spread the seeds through their feces, and the seeds are coated with a sticky substance that makes them adhere to farm equipment when the plants are mowed. It can form large patches that are difficult to eradicate due to their extensive root systems and sharp prickles. It is a major agricultural pest and a threat to native ecosystems.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf margins subentire, sinuate, or lobed, sinuses of lobes, when present, reaching less than 1/2 distance to midvein; apex of leaf lobes subacute to acute, sometimes rounded.
var. carolinense
1. Leaf margins deeply lobed, sinuses of lobes reaching more than 1/2 distance to midvein or almost to midvein; apex of leaf lobes typically rounded.
var. floridanum
Source FNA vol. 14. FNA vol. 14.
Parent taxa Solanaceae > Solanum Solanaceae > Solanum
Sibling taxa
S. americanum, S. aviculare, S. bahamense, S. campechiense, S. capsicoides, S. carolinense, S. chenopodioides, S. citrullifolium, S. cordicitum, S. davisense, S. deflexum, S. dimidiatum, S. diphyllum, S. donianum, S. douglasii, S. dulcamara, S. elaeagnifolium, S. emulans, S. erianthum, S. furcatum, S. hindsianum, S. interius, S. jamaicense, S. jamesii, S. laciniatum, S. lanceolatum, S. lumholtzianum, S. lycopersicum, S. marginatum, S. mauritianum, S. nigrescens, S. nigrum, S. nitidibaccatum, S. novomexicanum, S. perplexum, S. pseudocapsicum, S. pseudogracile, S. pumilum, S. rostratum, S. sarrachoides, S. seaforthianum, S. setigeroides, S. sisymbriifolium, S. stoloniferum, S. tampicense, S. tenuipes, S. torvum, S. triflorum, S. triquetrum, S. umbelliferum, S. wallacei
S. americanum, S. aviculare, S. bahamense, S. campechiense, S. capsicoides, S. chenopodioides, S. citrullifolium, S. cordicitum, S. davisense, S. deflexum, S. dimidiatum, S. diphyllum, S. donianum, S. douglasii, S. dulcamara, S. elaeagnifolium, S. emulans, S. erianthum, S. furcatum, S. hindsianum, S. interius, S. jamaicense, S. jamesii, S. laciniatum, S. lanceolatum, S. lumholtzianum, S. lycopersicum, S. marginatum, S. mauritianum, S. nigrescens, S. nigrum, S. nitidibaccatum, S. novomexicanum, S. perplexum, S. pseudocapsicum, S. pseudogracile, S. pumilum, S. rostratum, S. sarrachoides, S. seaforthianum, S. setigeroides, S. sisymbriifolium, S. stoloniferum, S. tampicense, S. tenuipes, S. torvum, S. triflorum, S. triquetrum, S. umbelliferum, S. viarum, S. wallacei
Subordinate taxa
S. carolinense var. carolinense, S. carolinense var. floridanum
Name authority Dunal in A. P. de Candolle and A. L. P. P. de Candolle: Prodr. 13(1): 240. (1852) Linnaeus: Sp. Pl. 1: 187. (1753)
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