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cut-leaf nightshade, three flower nightshade

blue nightshade, blue witch, blue witch or chaparral nightshade, bluewitch nightshade

Habit Herbs, annual, decumbent to prostrate, unarmed, to 0.4 m, fleshy, nearly glabrous to moderately pubescent, hairs unbranched, to 2 mm, eglandular, rarely glandular. Shrubs or subshrubs, erect or somewhat spreading, unarmed, to 1.5 m, glabrous to densely pubescent, hairs unbranched, to 2 mm, glandular or eglandular and dendritic.
Leaves

petiolate;

petiole 0.5–2.5 cm;

blade simple, elliptic to oblong, 2–5 × 1–3 cm, margins shallowly lobed to deeply and regularly pinnatifid with 3–6 lobes per side, lobe margins entire or occasionally coarsely lobed, base cuneate and decurrent.

petiolate;

petiole 0.2–1.5(–3) cm;

blade simple, lanceolate to ovate or obovate, (0.5–)1–4(–9) × 0.5–2(–6.5) cm, margins entire to pinnatifid with 1(–3) pairs of lobes at base, lobe margins entire to undulate, base attenuate to truncate, occasionally subcordate.

Inflorescences

extra-axillary, unbranched, umbel-like, 1–6-flowered, 1–3 cm.

terminal or lateral, leaf-opposed or extra-axillary, simple or once-branched, 5–20-flowered, 1–8 cm.

Pedicels

spreading and 0.5–1.5 cm in flower, reflexed and 0.5–1.5 cm in fruit.

inserted into small sleeve on inflorescence axis, 0.5–1.5 cm in flower, 1.2–2 cm in fruit.

Flowers

radially symmetric;

calyx accrescent and covering base of berry, unarmed, 2–4(–7) mm, moderately pubescent, lobes deltate, reflexed;

corolla white or light purple with green or purplish central star, stellate, 0.5–1 cm diam., with sparse interpetalar tissue;

stamens equal;

anthers narrowly ellipsoidal, 2.5–4 mm, dehiscent by terminal pores that open into longitudinal slits;

ovary glabrous.

radially symmetric;

calyx slightly accrescent, unarmed, 2.5–5.5 mm, glabrous to densely pubescent, hairs unbranched or dendritic, lobes broadly deltate;

corolla pale to deep purple or occasionally white, with green spots edged with white at base of lobes, spots separate or confluent, rotate, (1–)1.3–2.5 cm diam., with abundant interpetalar tissue;

stamens equal;

anthers ellipsoidal, slightly tapered, 3.5–4.5 mm, dehiscent by terminal pores that open into longitudinal slits;

ovary glabrous.

Berries

shiny dark green to purplish black, globose, 0.8–2 cm diam., glabrous, with 13–30 sclerotic granules.

green, greenish black, or black, globose, 1–2 cm diam., glabrous, without sclerotic granules.

Seeds

yellow, plump, 2–3 × 2–2.5 mm, minutely pitted.

reddish brown, flattened, ca. 2 × 1.5 mm, minutely pitted.

2n

= 24.

Solanum triflorum

Solanum umbelliferum

Phenology Flowering Apr–Sep. Flowering Feb–Nov (most of the year in California).
Habitat Disturbed areas, roadsides, stream banks, along railroad tracks, prairie dog towns. Sand dunes, chaparral, coastal sage scrub, rocky slopes, pine forests.
Elevation (0–)700–2900 m. ((0–)2300–9500 ft.) 0–2000 m. (0–6600 ft.)
Distribution
from FNA
AZ; CA; CO; IA; ID; KS; MA; MI; MN; MO; MT; ND; NE; NM; NV; OK; OR; SD; TX; UT; WA; WY; AB; BC; MB; SK; South America (Argentina) [Introduced in Europe, Africa, Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NV; OR; WA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
Discussion

Solanum triflorum is found in South America (Argentina) and is also considered to be native to central and western North America. It is occasionally adventive in the eastern United States. It is poisonous to livestock and can become a serious weed in cultivated fields, especially in the Great Plains.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Solanum umbelliferum is common in the western part of North America from Washington to Baja California. It is found throughout California except for Modoc Plateau, Desert Province, and Central Valley.

Past treatments have divided Solanum umbelliferum into a number of taxa based on leaf size and shape and pubescence type and density, but the most recent monograph (S. Knapp 2013) regarded it as one highly variable and widespread species in which no character discontinuities can be seen. Glabrous populations from northern California have been called S. parishii, sticky-glandular populations from central and southern California S. xanti, glabrous populations from southern California have been called var. glabrescens, and densely pubescent eglandular populations from central California have been called S. californicum Dunal. Island populations with larger leaves have been called S. clokeyi (but see 52. S. wallacei, a distinct endemic on Santa Catalina Island). A number of new varietal combinations were published by D. J. Keil (2018) to accommodate much of this regional and local variation, but the group needs thorough study using both molecular and morphological analysis across its range to assess the taxonomic validity of these segregants. Some of the variation may be environmental.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 14. FNA vol. 14.
Parent taxa Solanaceae > Solanum Solanaceae > Solanum
Sibling taxa
S. americanum, S. aviculare, S. bahamense, S. campechiense, S. capsicoides, S. carolinense, S. chenopodioides, S. citrullifolium, S. cordicitum, S. davisense, S. deflexum, S. dimidiatum, S. diphyllum, S. donianum, S. douglasii, S. dulcamara, S. elaeagnifolium, S. emulans, S. erianthum, S. furcatum, S. hindsianum, S. interius, S. jamaicense, S. jamesii, S. laciniatum, S. lanceolatum, S. lumholtzianum, S. lycopersicum, S. marginatum, S. mauritianum, S. nigrescens, S. nigrum, S. nitidibaccatum, S. novomexicanum, S. perplexum, S. pseudocapsicum, S. pseudogracile, S. pumilum, S. rostratum, S. sarrachoides, S. seaforthianum, S. setigeroides, S. sisymbriifolium, S. stoloniferum, S. tampicense, S. tenuipes, S. torvum, S. triquetrum, S. umbelliferum, S. viarum, S. wallacei
S. americanum, S. aviculare, S. bahamense, S. campechiense, S. capsicoides, S. carolinense, S. chenopodioides, S. citrullifolium, S. cordicitum, S. davisense, S. deflexum, S. dimidiatum, S. diphyllum, S. donianum, S. douglasii, S. dulcamara, S. elaeagnifolium, S. emulans, S. erianthum, S. furcatum, S. hindsianum, S. interius, S. jamaicense, S. jamesii, S. laciniatum, S. lanceolatum, S. lumholtzianum, S. lycopersicum, S. marginatum, S. mauritianum, S. nigrescens, S. nigrum, S. nitidibaccatum, S. novomexicanum, S. perplexum, S. pseudocapsicum, S. pseudogracile, S. pumilum, S. rostratum, S. sarrachoides, S. seaforthianum, S. setigeroides, S. sisymbriifolium, S. stoloniferum, S. tampicense, S. tenuipes, S. torvum, S. triflorum, S. triquetrum, S. viarum, S. wallacei
Synonyms S. clokeyi, S. obispoense, S. parishii, S. tenuilobatum, S. umbelliferum var. clokeyi, S. umbelliferum var. glabrescens, S. umbelliferum var. hoffmannii, S. umbelliferum var. incanum, S. umbelliferum var. intermedium, S. umbelliferum var. montanum, S. umbelliferum var. obispoense, S. umbelliferum var. xanti, S. wallacei var. clokeyi, S. xanti, S. xanti var. glabrescens, S. xanti var. hoffmannii, S. xanti var. intermedium, S. xanti var. montanum
Name authority Nuttall: Gen. N. Amer. Pl. 1: 128. (1818) Eschscholtz: Mém. Acad. Imp. Sci. St. Pétersbourg Hist. Acad. 10: 283. (1826)
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