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Brazilian nightshade

blue nightshade, blue witch, blue witch or chaparral nightshade, bluewitch nightshade

Habit Shrubs or subshrubs, erect or somewhat spreading, unarmed, to 1.5 m, glabrous to densely pubescent, hairs unbranched, to 2 mm, glandular or eglandular and dendritic.
Leaves

petiolate;

petioles twining around supports, 1–4 cm;

blade simple to compound, elliptic to broadly ovate, (2–)3.5–10(–13) × (1–)2–9(–11) cm, margins entire to divided with up to 4 pairs of leaflets, leaflet margins entire, base truncate or slightly cordate.

petiolate;

petiole 0.2–1.5(–3) cm;

blade simple, lanceolate to ovate or obovate, (0.5–)1–4(–9) × 0.5–2(–6.5) cm, margins entire to pinnatifid with 1(–3) pairs of lobes at base, lobe margins entire to undulate, base attenuate to truncate, occasionally subcordate.

Inflorescences

terminal, becoming lateral, extra-axillary, much-branched, to 100+-flowered, to 25+ cm.

terminal or lateral, leaf-opposed or extra-axillary, simple or once-branched, 5–20-flowered, 1–8 cm.

Pedicels

inserted into small sleeve on inflorescence axis, 0.8–1.4 cm in flower and fruit.

inserted into small sleeve on inflorescence axis, 0.5–1.5 cm in flower, 1.2–2 cm in fruit.

Flowers

radially symmetric;

calyx not accrescent, unarmed, ca. 0.5 mm, nearly truncate, glabrous or sparsely pubescent on lobe tips;

corolla purple, stellate, 1–2.5 cm diam., with sparse interpetalar tissue;

stamens unequal due to unequal filaments;

anthers ellipsoidal, 2–3 mm, dehiscent by terminal pores;

ovary glabrous.

radially symmetric;

calyx slightly accrescent, unarmed, 2.5–5.5 mm, glabrous to densely pubescent, hairs unbranched or dendritic, lobes broadly deltate;

corolla pale to deep purple or occasionally white, with green spots edged with white at base of lobes, spots separate or confluent, rotate, (1–)1.3–2.5 cm diam., with abundant interpetalar tissue;

stamens equal;

anthers ellipsoidal, slightly tapered, 3.5–4.5 mm, dehiscent by terminal pores that open into longitudinal slits;

ovary glabrous.

Berries

bright shiny red, globose, 0.8–1.5 cm diam., glabrous, without sclerotic granules.

green, greenish black, or black, globose, 1–2 cm diam., glabrous, without sclerotic granules.

Seeds

pale yellowish tan, flattened, 4–4.5 × 2.5-3 mm, minutely pitted.

reddish brown, flattened, ca. 2 × 1.5 mm, minutely pitted.

Climbing

or scrambling vines, woody, unarmed, to ca. 3 m, glabrous or sparsely pubescent, hairs white, unbranched, ca. 0.2 mm.

2n

= 24.

Solanum seaforthianum

Solanum umbelliferum

Phenology Flowering year-round. Flowering Feb–Nov (most of the year in California).
Habitat Disturbed sites. Sand dunes, chaparral, coastal sage scrub, rocky slopes, pine forests.
Elevation 0–200 m. (0–700 ft.) 0–2000 m. (0–6600 ft.)
Distribution
from FNA
FL; Mexico; West Indies; Central America (Belize, Costa Rica, Guatemala, Honduras, Nicaragua, Panama); South America (Brazil, Colombia, Ecuador, Venezuela) [Introduced in North America; introduced also elsewhere in South America (Argentina, Paraguay, Peru), Asia, Africa, Pacific Islands, Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NV; OR; WA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
Discussion

Solanum seaforthianum is widely cultivated as an ornamental and sporadically escapes in Florida. A similar species with twining petioles, S. laxum Sprengel, is occasionally cultivated in California. It is distinguished from S. seaforthianum by having tufts of hairs in the vein axils of the abaxial leaf surfaces, white rather than violet corollas, and equal stamens.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Solanum umbelliferum is common in the western part of North America from Washington to Baja California. It is found throughout California except for Modoc Plateau, Desert Province, and Central Valley.

Past treatments have divided Solanum umbelliferum into a number of taxa based on leaf size and shape and pubescence type and density, but the most recent monograph (S. Knapp 2013) regarded it as one highly variable and widespread species in which no character discontinuities can be seen. Glabrous populations from northern California have been called S. parishii, sticky-glandular populations from central and southern California S. xanti, glabrous populations from southern California have been called var. glabrescens, and densely pubescent eglandular populations from central California have been called S. californicum Dunal. Island populations with larger leaves have been called S. clokeyi (but see 52. S. wallacei, a distinct endemic on Santa Catalina Island). A number of new varietal combinations were published by D. J. Keil (2018) to accommodate much of this regional and local variation, but the group needs thorough study using both molecular and morphological analysis across its range to assess the taxonomic validity of these segregants. Some of the variation may be environmental.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 14. FNA vol. 14.
Parent taxa Solanaceae > Solanum Solanaceae > Solanum
Sibling taxa
S. americanum, S. aviculare, S. bahamense, S. campechiense, S. capsicoides, S. carolinense, S. chenopodioides, S. citrullifolium, S. cordicitum, S. davisense, S. deflexum, S. dimidiatum, S. diphyllum, S. donianum, S. douglasii, S. dulcamara, S. elaeagnifolium, S. emulans, S. erianthum, S. furcatum, S. hindsianum, S. interius, S. jamaicense, S. jamesii, S. laciniatum, S. lanceolatum, S. lumholtzianum, S. lycopersicum, S. marginatum, S. mauritianum, S. nigrescens, S. nigrum, S. nitidibaccatum, S. novomexicanum, S. perplexum, S. pseudocapsicum, S. pseudogracile, S. pumilum, S. rostratum, S. sarrachoides, S. setigeroides, S. sisymbriifolium, S. stoloniferum, S. tampicense, S. tenuipes, S. torvum, S. triflorum, S. triquetrum, S. umbelliferum, S. viarum, S. wallacei
S. americanum, S. aviculare, S. bahamense, S. campechiense, S. capsicoides, S. carolinense, S. chenopodioides, S. citrullifolium, S. cordicitum, S. davisense, S. deflexum, S. dimidiatum, S. diphyllum, S. donianum, S. douglasii, S. dulcamara, S. elaeagnifolium, S. emulans, S. erianthum, S. furcatum, S. hindsianum, S. interius, S. jamaicense, S. jamesii, S. laciniatum, S. lanceolatum, S. lumholtzianum, S. lycopersicum, S. marginatum, S. mauritianum, S. nigrescens, S. nigrum, S. nitidibaccatum, S. novomexicanum, S. perplexum, S. pseudocapsicum, S. pseudogracile, S. pumilum, S. rostratum, S. sarrachoides, S. seaforthianum, S. setigeroides, S. sisymbriifolium, S. stoloniferum, S. tampicense, S. tenuipes, S. torvum, S. triflorum, S. triquetrum, S. viarum, S. wallacei
Synonyms S. clokeyi, S. obispoense, S. parishii, S. tenuilobatum, S. umbelliferum var. clokeyi, S. umbelliferum var. glabrescens, S. umbelliferum var. hoffmannii, S. umbelliferum var. incanum, S. umbelliferum var. intermedium, S. umbelliferum var. montanum, S. umbelliferum var. obispoense, S. umbelliferum var. xanti, S. wallacei var. clokeyi, S. xanti, S. xanti var. glabrescens, S. xanti var. hoffmannii, S. xanti var. intermedium, S. xanti var. montanum
Name authority Andrews: Bot. Repos. 8: plate 504. (1808) Eschscholtz: Mém. Acad. Imp. Sci. St. Pétersbourg Hist. Acad. 10: 283. (1826)
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