Solanum nigrescens
Solanum umbelliferum
divine nightshade
blue nightshade, blue witch, blue witch or chaparral nightshade, bluewitch nightshade
petiolate;
petiole 0.5–2 cm;
blade simple, ovate to ovate-elliptic, 4–10.5 × 2–5 cm, margins entire or shallowly sinuate-dentate, base decurrent.
petiolate;
petiole 0.2–1.5(–3) cm;
blade simple, lanceolate to ovate or obovate, (0.5–)1–4(–9) × 0.5–2(–6.5) cm, margins entire to pinnatifid with 1(–3) pairs of lobes at base, lobe margins entire to undulate, base attenuate to truncate, occasionally subcordate.
extra-axillary or leaf-opposed, unbranched, umbel-like to racemelike, (2–)5–10-flowered, 1–3.5 cm.
terminal or lateral, leaf-opposed or extra-axillary, simple or once-branched, 5–20-flowered, 1–8 cm.
spreading and 0.5–1 cm in flower, spreading and 1–1.5 cm in fruit.
inserted into small sleeve on inflorescence axis, 0.5–1.5 cm in flower, 1.2–2 cm in fruit.
radially symmetric;
calyx not accrescent, unarmed, 1–2 mm, sparsely pubescent, lobes deltate;
corolla white, rarely purplish, often with green or purplish central star, stellate, 1–1.5 cm diam., with sparse interpetalar tissue;
stamens equal;
anthers ellipsoidal, 2–3 mm, dehiscent by terminal pores that open into longitudinal slits;
ovary glabrous.
radially symmetric;
calyx slightly accrescent, unarmed, 2.5–5.5 mm, glabrous to densely pubescent, hairs unbranched or dendritic, lobes broadly deltate;
corolla pale to deep purple or occasionally white, with green spots edged with white at base of lobes, spots separate or confluent, rotate, (1–)1.3–2.5 cm diam., with abundant interpetalar tissue;
stamens equal;
anthers ellipsoidal, slightly tapered, 3.5–4.5 mm, dehiscent by terminal pores that open into longitudinal slits;
ovary glabrous.
dull green or purplish, globose, 0.5–0.8 cm diam., glabrous, with (4–)5–6(–13) sclerotic granules.
green, greenish black, or black, globose, 1–2 cm diam., glabrous, without sclerotic granules.
tan, flattened, 1.2–1.5 × 1–1.1 mm, finely pitted.
reddish brown, flattened, ca. 2 × 1.5 mm, minutely pitted.
= 24.
Solanum nigrescens
Solanum umbelliferum
Solanum nigrescens is widespread in Central and northern South America and the Caribbean and extends northward into the southeastern United States along the Gulf Coast and slightly inland. Where sympatric with S. americanum, it can be distinguished by its longer anthers and dull green or purplish berries with appressed to spreading calyx lobes. Plants collected as weeds in rice and sugarcane fields of Louisiana and provisionally identified as the Chinese species S. merrillianum T. N. Liou are somewhat intermediate between S. americanum and S. nigrescens and could represent recent hybrid populations (S. Knapp et al. 2019). Solanum nigrescens differs from S. douglasii in its shorter anthers and longer filaments relative to anther length; moreover, S. douglasii is usually found west of the Rocky Mountains, whereas S. nigrescens occurs in the southeastern United States. The ranges of S. nigrescens and S. interius overlap (for example, in Texas). Solanum nigrescens may be distinguished from S. interius by its usually acute calyx lobes, smaller seeds, and more numerous sclerotic granules in the fruits. Solanum nigrescens differs from S. nigrum in its more slender peduncles and pedicels, smaller seeds, and fruits with sclerotic granules.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Solanum umbelliferum is common in the western part of North America from Washington to Baja California. It is found throughout California except for Modoc Plateau, Desert Province, and Central Valley.
Past treatments have divided Solanum umbelliferum into a number of taxa based on leaf size and shape and pubescence type and density, but the most recent monograph (S. Knapp 2013) regarded it as one highly variable and widespread species in which no character discontinuities can be seen. Glabrous populations from northern California have been called S. parishii, sticky-glandular populations from central and southern California S. xanti, glabrous populations from southern California have been called var. glabrescens, and densely pubescent eglandular populations from central California have been called S. californicum Dunal. Island populations with larger leaves have been called S. clokeyi (but see 52. S. wallacei, a distinct endemic on Santa Catalina Island). A number of new varietal combinations were published by D. J. Keil (2018) to accommodate much of this regional and local variation, but the group needs thorough study using both molecular and morphological analysis across its range to assess the taxonomic validity of these segregants. Some of the variation may be environmental.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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