Solanum nigrescens |
Solanum dulcamara |
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divine nightshade |
bitter nightshade, bittersweet, bittersweet nightshade, blue bindweed, climbing nightshade, European bittersweet, felonwort |
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Habit | Herbs, perennial, erect to somewhat sprawling, unarmed, to 3 m, nearly glabrous to moderately pubescent, hairs unbranched, to 1 mm, eglandular. | |
Leaves | petiolate; petiole 0.5–2 cm; blade simple, ovate to ovate-elliptic, 4–10.5 × 2–5 cm, margins entire or shallowly sinuate-dentate, base decurrent. |
petiolate; petiole 0.5–5 cm; blade simple, elliptic or ovate to cordate, 2.5–12 × 1.2–9 cm, margins entire to deeply pinnatifid and usually 3-lobed near base, lobe margins entire, base truncate to cordate. |
Inflorescences | extra-axillary or leaf-opposed, unbranched, umbel-like to racemelike, (2–)5–10-flowered, 1–3.5 cm. |
terminal or lateral, extra-axillary, much-branched, 7–40-flowered, (1–)4–15 cm. |
Pedicels | spreading and 0.5–1 cm in flower, spreading and 1–1.5 cm in fruit. |
inserted into a small sleeve on the inflorescence axis, 0.6–1.2 cm in flower and fruit. |
Flowers | radially symmetric; calyx not accrescent, unarmed, 1–2 mm, sparsely pubescent, lobes deltate; corolla white, rarely purplish, often with green or purplish central star, stellate, 1–1.5 cm diam., with sparse interpetalar tissue; stamens equal; anthers ellipsoidal, 2–3 mm, dehiscent by terminal pores that open into longitudinal slits; ovary glabrous. |
radially symmetric; calyx not accrescent, unarmed, 1–2 mm, glabrous to densely pubescent, lobes triangular, shallow; corolla purple (rarely white), with green and white shiny spots at base of each lobe, deeply stellate, 1.5–2 cm diam., without interpetalar tissue; stamens equal; anthers oblong, slightly tapered, 4.5–6 mm, dehiscent by terminal pores that often open into longitudinal slits; ovary glabrous. |
Berries | dull green or purplish, globose, 0.5–0.8 cm diam., glabrous, with (4–)5–6(–13) sclerotic granules. |
bright shiny red, globose to ellipsoidal, 0.5–1.5 × 0.5–1 cm, glabrous, without sclerotic granules. |
Seeds | tan, flattened, 1.2–1.5 × 1–1.1 mm, finely pitted. |
pale yellow or tan, flattened, 2–3 mm diam., minutely pitted. |
Vines | , climbing or scrambling, herbaceous or woody, unarmed, to 8–10 m, sparsely to densely pubescent, hairs unbranched and/or dendritic, rarely glabrous. |
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2n | = 24. |
= 24. |
Solanum nigrescens |
Solanum dulcamara |
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Phenology | Flowering year-round. | Flowering May–Nov. |
Habitat | Deciduous and coniferous forests, fields, swampy areas. | Weedy, in a wide variety of habitats, often associated with water. |
Elevation | 0–1500 m. (0–4900 ft.) | 0–2000 m. (0–6600 ft.) |
Distribution |
AL; FL; LA; MS; NC; NM; TX; Mexico; Central America; South America; West Indies
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AL; AZ; CA; CO; CT; DC; FL; GA; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; RI; SD; TN; UT; VA; VT; WA; WI; WV; WY; BC; MB; NB; NF; NS; ON; PE; QC; SK; Eurasia
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Discussion | Solanum nigrescens is widespread in Central and northern South America and the Caribbean and extends northward into the southeastern United States along the Gulf Coast and slightly inland. Where sympatric with S. americanum, it can be distinguished by its longer anthers and dull green or purplish berries with appressed to spreading calyx lobes. Plants collected as weeds in rice and sugarcane fields of Louisiana and provisionally identified as the Chinese species S. merrillianum T. N. Liou are somewhat intermediate between S. americanum and S. nigrescens and could represent recent hybrid populations (S. Knapp et al. 2019). Solanum nigrescens differs from S. douglasii in its shorter anthers and longer filaments relative to anther length; moreover, S. douglasii is usually found west of the Rocky Mountains, whereas S. nigrescens occurs in the southeastern United States. The ranges of S. nigrescens and S. interius overlap (for example, in Texas). Solanum nigrescens may be distinguished from S. interius by its usually acute calyx lobes, smaller seeds, and more numerous sclerotic granules in the fruits. Solanum nigrescens differs from S. nigrum in its more slender peduncles and pedicels, smaller seeds, and fruits with sclerotic granules. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Solanum dulcamara is widely distributed across Eurasia and boreal North America. The North American populations are thought to be introductions, but it is possible that the species has a truly circumboreal distribution. A white-flowered form has been recognized by some authors as S. dulcamara forma albiflorum House and an especially pubescent form as S. dulcamara var. villosissimum, but variation in a number of morphological features is continuous across the range of the species and these and other variants are not recognized in the latest monograph of the group (S. Knapp 2013). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 14. | FNA vol. 14. |
Parent taxa | Solanaceae > Solanum | Solanaceae > Solanum |
Sibling taxa | ||
Synonyms | S. dulcamara var. villosissimum | |
Name authority | M. Martens & Galeotti: Bull. Acad. Roy. Sci. Bruxelles 12(1): 140. (1845) | Linnaeus: Sp. Pl. 1: 185. (1753) |
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