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earleaf nightshade

blue nightshade, blue witch, blue witch or chaparral nightshade, bluewitch nightshade

Habit Shrubs or small trees, erect, unarmed, 2–12 m, densely pubescent, hairs white, sessile to long-stalked, stellate to echinoid. Shrubs or subshrubs, erect or somewhat spreading, unarmed, to 1.5 m, glabrous to densely pubescent, hairs unbranched, to 2 mm, glandular or eglandular and dendritic.
Leaves

petiolate;

petiole 1–8 cm;

blade simple, elliptic to ovate, 11–31 × 4–14 cm, margins entire, base acute, often with smaller axillary leaves.

petiolate;

petiole 0.2–1.5(–3) cm;

blade simple, lanceolate to ovate or obovate, (0.5–)1–4(–9) × 0.5–2(–6.5) cm, margins entire to pinnatifid with 1(–3) pairs of lobes at base, lobe margins entire to undulate, base attenuate to truncate, occasionally subcordate.

Inflorescences

terminal, becoming leaf-opposed, much-branched, 50–100-flowered, 5–24 cm.

terminal or lateral, leaf-opposed or extra-axillary, simple or once-branched, 5–20-flowered, 1–8 cm.

Pedicels

erect and 0.2–0.5 cm in flower and fruit.

inserted into small sleeve on inflorescence axis, 0.5–1.5 cm in flower, 1.2–2 cm in fruit.

Flowers

radially symmetric;

calyx slightly accrescent, unarmed, 4–7.5 mm, densely pubescent, lobes deltate;

corolla purple, stellate-pentagonal, 1–1.5 cm diam., with abundant interpetalar tissue;

stamens equal;

anthers oblong, 2–3.5 mm, dehiscent by terminal pores that open into longitudinal slits;

ovary tomentose.

radially symmetric;

calyx slightly accrescent, unarmed, 2.5–5.5 mm, glabrous to densely pubescent, hairs unbranched or dendritic, lobes broadly deltate;

corolla pale to deep purple or occasionally white, with green spots edged with white at base of lobes, spots separate or confluent, rotate, (1–)1.3–2.5 cm diam., with abundant interpetalar tissue;

stamens equal;

anthers ellipsoidal, slightly tapered, 3.5–4.5 mm, dehiscent by terminal pores that open into longitudinal slits;

ovary glabrous.

Berries

yellow, globose, 1–1.5 cm diam., tomentose, without sclerotic granules.

green, greenish black, or black, globose, 1–2 cm diam., glabrous, without sclerotic granules.

Seeds

yellowish brown, flattened, 1.5–2.5 × 1.5–2 mm, minutely pitted.

reddish brown, flattened, ca. 2 × 1.5 mm, minutely pitted.

2n

= 24.

Solanum mauritianum

Solanum umbelliferum

Phenology Flowering Mar–Jul. Flowering Feb–Nov (most of the year in California).
Habitat Disturbed sites. Sand dunes, chaparral, coastal sage scrub, rocky slopes, pine forests.
Elevation 0–500 m. (0–1600 ft.) 0–2000 m. (0–6600 ft.)
Distribution
from FNA
CA; FL; South America (Brazil, Uruguay) [Introduced also in Asia (India), Africa, Atlantic Islands, Indian Ocean Islands, Pacific Islands, Australia]
[BONAP county map]
from FNA
AZ; CA; NV; OR; WA; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
Discussion

In Florida, Solanum mauritianum has become naturalized and common only at one site in Pasco County. It also occurs frequently in southern California from Santa Barbara south to San Diego with urban waifs in the Bay Area. It appears to be spreading into relatively undisturbed riparian areas in the San Gabriel Mountains and may become a widespread pest.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Solanum umbelliferum is common in the western part of North America from Washington to Baja California. It is found throughout California except for Modoc Plateau, Desert Province, and Central Valley.

Past treatments have divided Solanum umbelliferum into a number of taxa based on leaf size and shape and pubescence type and density, but the most recent monograph (S. Knapp 2013) regarded it as one highly variable and widespread species in which no character discontinuities can be seen. Glabrous populations from northern California have been called S. parishii, sticky-glandular populations from central and southern California S. xanti, glabrous populations from southern California have been called var. glabrescens, and densely pubescent eglandular populations from central California have been called S. californicum Dunal. Island populations with larger leaves have been called S. clokeyi (but see 52. S. wallacei, a distinct endemic on Santa Catalina Island). A number of new varietal combinations were published by D. J. Keil (2018) to accommodate much of this regional and local variation, but the group needs thorough study using both molecular and morphological analysis across its range to assess the taxonomic validity of these segregants. Some of the variation may be environmental.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 14. FNA vol. 14.
Parent taxa Solanaceae > Solanum Solanaceae > Solanum
Sibling taxa
S. americanum, S. aviculare, S. bahamense, S. campechiense, S. capsicoides, S. carolinense, S. chenopodioides, S. citrullifolium, S. cordicitum, S. davisense, S. deflexum, S. dimidiatum, S. diphyllum, S. donianum, S. douglasii, S. dulcamara, S. elaeagnifolium, S. emulans, S. erianthum, S. furcatum, S. hindsianum, S. interius, S. jamaicense, S. jamesii, S. laciniatum, S. lanceolatum, S. lumholtzianum, S. lycopersicum, S. marginatum, S. nigrescens, S. nigrum, S. nitidibaccatum, S. novomexicanum, S. perplexum, S. pseudocapsicum, S. pseudogracile, S. pumilum, S. rostratum, S. sarrachoides, S. seaforthianum, S. setigeroides, S. sisymbriifolium, S. stoloniferum, S. tampicense, S. tenuipes, S. torvum, S. triflorum, S. triquetrum, S. umbelliferum, S. viarum, S. wallacei
S. americanum, S. aviculare, S. bahamense, S. campechiense, S. capsicoides, S. carolinense, S. chenopodioides, S. citrullifolium, S. cordicitum, S. davisense, S. deflexum, S. dimidiatum, S. diphyllum, S. donianum, S. douglasii, S. dulcamara, S. elaeagnifolium, S. emulans, S. erianthum, S. furcatum, S. hindsianum, S. interius, S. jamaicense, S. jamesii, S. laciniatum, S. lanceolatum, S. lumholtzianum, S. lycopersicum, S. marginatum, S. mauritianum, S. nigrescens, S. nigrum, S. nitidibaccatum, S. novomexicanum, S. perplexum, S. pseudocapsicum, S. pseudogracile, S. pumilum, S. rostratum, S. sarrachoides, S. seaforthianum, S. setigeroides, S. sisymbriifolium, S. stoloniferum, S. tampicense, S. tenuipes, S. torvum, S. triflorum, S. triquetrum, S. viarum, S. wallacei
Synonyms S. auriculatum S. clokeyi, S. obispoense, S. parishii, S. tenuilobatum, S. umbelliferum var. clokeyi, S. umbelliferum var. glabrescens, S. umbelliferum var. hoffmannii, S. umbelliferum var. incanum, S. umbelliferum var. intermedium, S. umbelliferum var. montanum, S. umbelliferum var. obispoense, S. umbelliferum var. xanti, S. wallacei var. clokeyi, S. xanti, S. xanti var. glabrescens, S. xanti var. hoffmannii, S. xanti var. intermedium, S. xanti var. montanum
Name authority Scopoli: Delic. Fl. Faun. Insubr. 3: 16, plate 8. (1788) Eschscholtz: Mém. Acad. Imp. Sci. St. Pétersbourg Hist. Acad. 10: 283. (1826)
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