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bitter nightshade, bittersweet, bittersweet nightshade, blue bindweed, climbing nightshade, European bittersweet, felonwort

Catalina nightshade, greasy or Santa Catalina or Wallace's nightshade, northern island nightshade, Wallace's nightshade

Habit Shrubs or small trees, erect to spreading, unarmed, 1–1.5(–3) m, densely pubescent, hairs transparent, unbranched, to ca. 3 mm, usually glandular.
Leaves

petiolate;

petiole 0.5–5 cm;

blade simple, elliptic or ovate to cordate, 2.5–12 × 1.2–9 cm, margins entire to deeply pinnatifid and usually 3-lobed near base, lobe margins entire, base truncate to cordate.

petiolate;

petiole 1–2.5(–4) cm;

blade simple, elliptic to obovate, 3–11(–14) × 1.6–5.5(–9) cm, margins entire or slightly undulate, occasionally with 2 small lobes at base, lobe margins entire, base truncate or acute.

Inflorescences

terminal or lateral, extra-axillary, much-branched, 7–40-flowered, (1–)4–15 cm.

terminal or lateral, leaf-opposed or extra-axillary, usually once-branched, occasionally more, 20–30-flowered, (2–)4–10 cm.

Pedicels

inserted into a small sleeve on the inflorescence axis, 0.6–1.2 cm in flower and fruit.

inserted into small sleeve on inflorescence axis, 1.5–2 cm in flower and fruit.

Flowers

radially symmetric;

calyx not accrescent, unarmed, 1–2 mm, glabrous to densely pubescent, lobes triangular, shallow;

corolla purple (rarely white), with green and white shiny spots at base of each lobe, deeply stellate, 1.5–2 cm diam., without interpetalar tissue;

stamens equal;

anthers oblong, slightly tapered, 4.5–6 mm, dehiscent by terminal pores that often open into longitudinal slits;

ovary glabrous.

radially symmetric;

calyx not accrescent, unarmed, 6–7 mm, densely pubescent, lobes deltate;

corolla violet to purple with or without green spots at base of lobes, spots usually small and not confluent, rotate, 3–4.5 cm diam., with abundant interpetalar tissue;

stamens equal;

anthers ellipsoidal, 4.5–5 mm, dehiscent by terminal pores that open into longitudinal slits;

ovary glabrous.

Berries

bright shiny red, globose to ellipsoidal, 0.5–1.5 × 0.5–1 cm, glabrous, without sclerotic granules.

shiny green, turning yellow then black, globose, 3–4 cm diam., glabrous, without sclerotic granules.

Seeds

pale yellow or tan, flattened, 2–3 mm diam., minutely pitted.

reddish brown, plump to flattened, 1.5–2 × 1–1.5 mm, minutely pitted.

Vines

, climbing or scrambling, herbaceous or woody, unarmed, to 8–10 m, sparsely to densely pubescent, hairs unbranched and/or dendritic, rarely glabrous.

2n

= 24.

Solanum dulcamara

Solanum wallacei

Phenology Flowering May–Nov. Flowering Mar–Aug.
Habitat Weedy, in a wide variety of habitats, often associated with water. Chaparral, open areas, canyon bottoms.
Elevation 0–2000 m. (0–6600 ft.) 0–300 m. (0–1000 ft.)
Distribution
from FNA
AL; AZ; CA; CO; CT; DC; FL; GA; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; RI; SD; TN; UT; VA; VT; WA; WI; WV; WY; BC; MB; NB; NF; NS; ON; PE; QC; SK; Eurasia
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Solanum dulcamara is widely distributed across Eurasia and boreal North America. The North American populations are thought to be introductions, but it is possible that the species has a truly circumboreal distribution. A white-flowered form has been recognized by some authors as S. dulcamara forma albiflorum House and an especially pubescent form as S. dulcamara var. villosissimum, but variation in a number of morphological features is continuous across the range of the species and these and other variants are not recognized in the latest monograph of the group (S. Knapp 2013).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Solanum wallacei is endemic to Santa Catalina Island.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 14. FNA vol. 14.
Parent taxa Solanaceae > Solanum Solanaceae > Solanum
Sibling taxa
S. americanum, S. aviculare, S. bahamense, S. campechiense, S. capsicoides, S. carolinense, S. chenopodioides, S. citrullifolium, S. cordicitum, S. davisense, S. deflexum, S. dimidiatum, S. diphyllum, S. donianum, S. douglasii, S. elaeagnifolium, S. emulans, S. erianthum, S. furcatum, S. hindsianum, S. interius, S. jamaicense, S. jamesii, S. laciniatum, S. lanceolatum, S. lumholtzianum, S. lycopersicum, S. marginatum, S. mauritianum, S. nigrescens, S. nigrum, S. nitidibaccatum, S. novomexicanum, S. perplexum, S. pseudocapsicum, S. pseudogracile, S. pumilum, S. rostratum, S. sarrachoides, S. seaforthianum, S. setigeroides, S. sisymbriifolium, S. stoloniferum, S. tampicense, S. tenuipes, S. torvum, S. triflorum, S. triquetrum, S. umbelliferum, S. viarum, S. wallacei
S. americanum, S. aviculare, S. bahamense, S. campechiense, S. capsicoides, S. carolinense, S. chenopodioides, S. citrullifolium, S. cordicitum, S. davisense, S. deflexum, S. dimidiatum, S. diphyllum, S. donianum, S. douglasii, S. dulcamara, S. elaeagnifolium, S. emulans, S. erianthum, S. furcatum, S. hindsianum, S. interius, S. jamaicense, S. jamesii, S. laciniatum, S. lanceolatum, S. lumholtzianum, S. lycopersicum, S. marginatum, S. mauritianum, S. nigrescens, S. nigrum, S. nitidibaccatum, S. novomexicanum, S. perplexum, S. pseudocapsicum, S. pseudogracile, S. pumilum, S. rostratum, S. sarrachoides, S. seaforthianum, S. setigeroides, S. sisymbriifolium, S. stoloniferum, S. tampicense, S. tenuipes, S. torvum, S. triflorum, S. triquetrum, S. umbelliferum, S. viarum
Synonyms S. dulcamara var. villosissimum S. xanti var. wallacei, S. umbelliferum var. wallacei
Name authority Linnaeus: Sp. Pl. 1: 185. (1753) (A. Gray) Parish: Proc. Calif. Acad. Sci., ser. 3, 2: 166. (1901)
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