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bitter nightshade, bittersweet, bittersweet nightshade, blue bindweed, climbing nightshade, European bittersweet, felonwort

fancy nightshade

Habit Herbs, perennial, spreading, moderately to densely armed, 0.2–0.5 m, prickles pale, straight, to 8 mm, glabrate to sparsely or moderately pubescent, hairs unbranched, glandular or eglandular, abaxial leaf surfaces with some sessile stellate hairs, 4–6-rayed, central ray equal to lateral rays.
Leaves

petiolate;

petiole 0.5–5 cm;

blade simple, elliptic or ovate to cordate, 2.5–12 × 1.2–9 cm, margins entire to deeply pinnatifid and usually 3-lobed near base, lobe margins entire, base truncate to cordate.

petiolate;

petiole 2–6 cm;

blade simple to 2–3 times lobed to compound, broadly ovate to elliptic, 4–9 × 2–7 cm, margins 2–3 times lobed to 2–3 times divided with 2–3 main leaflets per side, leaflets lobed to pinnately dissected, base truncate.

Inflorescences

terminal or lateral, extra-axillary, much-branched, 7–40-flowered, (1–)4–15 cm.

extra-axillary, unbranched, 6–9-flowered, 4–9 cm.

Pedicels

inserted into a small sleeve on the inflorescence axis, 0.6–1.2 cm in flower and fruit.

1–2 cm in flower, erect and 2–3 cm in fruit.

Flowers

radially symmetric;

calyx not accrescent, unarmed, 1–2 mm, glabrous to densely pubescent, lobes triangular, shallow;

corolla purple (rarely white), with green and white shiny spots at base of each lobe, deeply stellate, 1.5–2 cm diam., without interpetalar tissue;

stamens equal;

anthers oblong, slightly tapered, 4.5–6 mm, dehiscent by terminal pores that often open into longitudinal slits;

ovary glabrous.

bilaterally symmetric;

calyx accrescent and almost completely and tightly covering fruit, moderately to densely prickly, 4–6 mm, sparsely to moderately pubescent, lobes linear to lanceolate;

corolla violet or blue, stellate, 2.5–3.5 cm diam., with sparse interpetalar tissue;

stamens unequal, lowermost much longer and curved;

anthers narrow and tapered, dehiscent by terminal pores, short anthers 7–10 mm, longer anther 12–20 mm;

ovary glabrous.

Berries

bright shiny red, globose to ellipsoidal, 0.5–1.5 × 0.5–1 cm, glabrous, without sclerotic granules.

brown, globose to depressed-globose, 0.7–1 cm diam., glabrous, dry, without sclerotic granules.

Seeds

pale yellow or tan, flattened, 2–3 mm diam., minutely pitted.

dark brown, plump, 2.8–3.6 × 2–3 mm, minutely pitted and irregularly ridged.

Vines

, climbing or scrambling, herbaceous or woody, unarmed, to 8–10 m, sparsely to densely pubescent, hairs unbranched and/or dendritic, rarely glabrous.

2n

= 24.

= 24.

Solanum dulcamara

Solanum tenuipes

Phenology Flowering May–Nov.
Habitat Weedy, in a wide variety of habitats, often associated with water.
Elevation 0–2000 m. (0–6600 ft.)
Distribution
from FNA
AL; AZ; CA; CO; CT; DC; FL; GA; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; RI; SD; TN; UT; VA; VT; WA; WI; WV; WY; BC; MB; NB; NF; NS; ON; PE; QC; SK; Eurasia
[WildflowerSearch map]
[BONAP county map]
from USDA
nc Mexico; Texas
[BONAP county map]
Discussion

Solanum dulcamara is widely distributed across Eurasia and boreal North America. The North American populations are thought to be introductions, but it is possible that the species has a truly circumboreal distribution. A white-flowered form has been recognized by some authors as S. dulcamara forma albiflorum House and an especially pubescent form as S. dulcamara var. villosissimum, but variation in a number of morphological features is continuous across the range of the species and these and other variants are not recognized in the latest monograph of the group (S. Knapp 2013).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Larger leaves 3 times pinnatifid; seeds 3.1–3.6 mm.
var. tenuipes
1. Leaves usually only 2 times pinnatifid; seeds to 3 mm.
var. latisectum
Source FNA vol. 14. FNA vol. 14.
Parent taxa Solanaceae > Solanum Solanaceae > Solanum
Sibling taxa
S. americanum, S. aviculare, S. bahamense, S. campechiense, S. capsicoides, S. carolinense, S. chenopodioides, S. citrullifolium, S. cordicitum, S. davisense, S. deflexum, S. dimidiatum, S. diphyllum, S. donianum, S. douglasii, S. elaeagnifolium, S. emulans, S. erianthum, S. furcatum, S. hindsianum, S. interius, S. jamaicense, S. jamesii, S. laciniatum, S. lanceolatum, S. lumholtzianum, S. lycopersicum, S. marginatum, S. mauritianum, S. nigrescens, S. nigrum, S. nitidibaccatum, S. novomexicanum, S. perplexum, S. pseudocapsicum, S. pseudogracile, S. pumilum, S. rostratum, S. sarrachoides, S. seaforthianum, S. setigeroides, S. sisymbriifolium, S. stoloniferum, S. tampicense, S. tenuipes, S. torvum, S. triflorum, S. triquetrum, S. umbelliferum, S. viarum, S. wallacei
S. americanum, S. aviculare, S. bahamense, S. campechiense, S. capsicoides, S. carolinense, S. chenopodioides, S. citrullifolium, S. cordicitum, S. davisense, S. deflexum, S. dimidiatum, S. diphyllum, S. donianum, S. douglasii, S. dulcamara, S. elaeagnifolium, S. emulans, S. erianthum, S. furcatum, S. hindsianum, S. interius, S. jamaicense, S. jamesii, S. laciniatum, S. lanceolatum, S. lumholtzianum, S. lycopersicum, S. marginatum, S. mauritianum, S. nigrescens, S. nigrum, S. nitidibaccatum, S. novomexicanum, S. perplexum, S. pseudocapsicum, S. pseudogracile, S. pumilum, S. rostratum, S. sarrachoides, S. seaforthianum, S. setigeroides, S. sisymbriifolium, S. stoloniferum, S. tampicense, S. torvum, S. triflorum, S. triquetrum, S. umbelliferum, S. viarum, S. wallacei
Subordinate taxa
S. tenuipes var. latisectum, S. tenuipes var. tenuipes
Synonyms S. dulcamara var. villosissimum
Name authority Linnaeus: Sp. Pl. 1: 185. (1753) Bartlett: Proc. Amer. Acad. Arts 44: 629. (1909)
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