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bitter nightshade, bittersweet, bittersweet nightshade, blue bindweed, climbing nightshade, European bittersweet, felonwort

coast nightshade, fork nightshade

Habit Herbs, annual or perennial, erect to sprawling, unarmed, to 1 m, sparsely pubescent, hairs unbranched, to 0.5 mm, eglandular.
Leaves

petiolate;

petiole 0.5–5 cm;

blade simple, elliptic or ovate to cordate, 2.5–12 × 1.2–9 cm, margins entire to deeply pinnatifid and usually 3-lobed near base, lobe margins entire, base truncate to cordate.

petiolate;

petiole 1–3.5 cm;

blade simple, ovate-lanceolate, 3–10 × 2–5 cm, margins entire to sinuate-dentate, base cuneate to truncate.

Inflorescences

terminal or lateral, extra-axillary, much-branched, 7–40-flowered, (1–)4–15 cm.

extra-axillary, forked, umbel-like or racemelike, 6–14-flowered, 1.5–3 cm.

Pedicels

inserted into a small sleeve on the inflorescence axis, 0.6–1.2 cm in flower and fruit.

straight and spreading and 0.5–1 cm in flower, strongly reflexed and 0.5–1 cm in fruit.

Flowers

radially symmetric;

calyx not accrescent, unarmed, 1–2 mm, glabrous to densely pubescent, lobes triangular, shallow;

corolla purple (rarely white), with green and white shiny spots at base of each lobe, deeply stellate, 1.5–2 cm diam., without interpetalar tissue;

stamens equal;

anthers oblong, slightly tapered, 4.5–6 mm, dehiscent by terminal pores that often open into longitudinal slits;

ovary glabrous.

radially symmetric;

calyx not accrescent, unarmed, 3–4 mm, sparsely pubescent, lobes obtuse;

corolla white to pale purple with yellowish or greenish central star, stellate, 1–2 cm diam., with sparse interpetalar tissue;

stamens equal;

anthers ellipsoidal, 2.5–3.5 mm, dehiscent by terminal pores that open into longitudinal slits;

ovary glabrous.

Berries

bright shiny red, globose to ellipsoidal, 0.5–1.5 × 0.5–1 cm, glabrous, without sclerotic granules.

dull green to purple, globose, 0.5–0.9 cm diam., glabrous, with 6–14 sclerotic granules per fruit.

Seeds

pale yellow or tan, flattened, 2–3 mm diam., minutely pitted.

pale yellow to light brown, flattened, 1.5–2 × 1–1.5 mm, finely reticulate.

Vines

, climbing or scrambling, herbaceous or woody, unarmed, to 8–10 m, sparsely to densely pubescent, hairs unbranched and/or dendritic, rarely glabrous.

2n

= 24.

= 72.

Solanum dulcamara

Solanum furcatum

Phenology Flowering May–Nov. Flowering May–Oct.
Habitat Weedy, in a wide variety of habitats, often associated with water. Open and disturbed areas near sea cliffs, bluffs, and on sand dunes.
Elevation 0–2000 m. (0–6600 ft.) 0–500 m. (0–1600 ft.)
Distribution
from FNA
AL; AZ; CA; CO; CT; DC; FL; GA; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; RI; SD; TN; UT; VA; VT; WA; WI; WV; WY; BC; MB; NB; NF; NS; ON; PE; QC; SK; Eurasia
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR; WA; South America (Argentina, Chile) [Introduced also in Pacific Islands (New Zealand), Australia]
[WildflowerSearch map]
[BONAP county map]
Discussion

Solanum dulcamara is widely distributed across Eurasia and boreal North America. The North American populations are thought to be introductions, but it is possible that the species has a truly circumboreal distribution. A white-flowered form has been recognized by some authors as S. dulcamara forma albiflorum House and an especially pubescent form as S. dulcamara var. villosissimum, but variation in a number of morphological features is continuous across the range of the species and these and other variants are not recognized in the latest monograph of the group (S. Knapp 2013).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Solanum furcatum is found in coastal environments in the western United States. M. Nee (1993) stated that the name S. gayanum (J. Remy) F. Philippi has been misapplied to plants of S. furcatum, but no basis can be found for this assertion and the two species are morphologically very different. Solanum gayanum, a synonym of S. crispum Ruiz & Pavon, and native to Chile, is cultivated and perhaps naturalized in San Francisco, California (P. A. Munz 1968).

Solanum furcatum can be distinguished from the similar and sympatric S. douglasii by its usually forked inflorescences and fruits with usually more than ten sclerotic granules. A distinctive character of S. furcatum is the long style that is about twice the length of the anthers.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 14. FNA vol. 14.
Parent taxa Solanaceae > Solanum Solanaceae > Solanum
Sibling taxa
S. americanum, S. aviculare, S. bahamense, S. campechiense, S. capsicoides, S. carolinense, S. chenopodioides, S. citrullifolium, S. cordicitum, S. davisense, S. deflexum, S. dimidiatum, S. diphyllum, S. donianum, S. douglasii, S. elaeagnifolium, S. emulans, S. erianthum, S. furcatum, S. hindsianum, S. interius, S. jamaicense, S. jamesii, S. laciniatum, S. lanceolatum, S. lumholtzianum, S. lycopersicum, S. marginatum, S. mauritianum, S. nigrescens, S. nigrum, S. nitidibaccatum, S. novomexicanum, S. perplexum, S. pseudocapsicum, S. pseudogracile, S. pumilum, S. rostratum, S. sarrachoides, S. seaforthianum, S. setigeroides, S. sisymbriifolium, S. stoloniferum, S. tampicense, S. tenuipes, S. torvum, S. triflorum, S. triquetrum, S. umbelliferum, S. viarum, S. wallacei
S. americanum, S. aviculare, S. bahamense, S. campechiense, S. capsicoides, S. carolinense, S. chenopodioides, S. citrullifolium, S. cordicitum, S. davisense, S. deflexum, S. dimidiatum, S. diphyllum, S. donianum, S. douglasii, S. dulcamara, S. elaeagnifolium, S. emulans, S. erianthum, S. hindsianum, S. interius, S. jamaicense, S. jamesii, S. laciniatum, S. lanceolatum, S. lumholtzianum, S. lycopersicum, S. marginatum, S. mauritianum, S. nigrescens, S. nigrum, S. nitidibaccatum, S. novomexicanum, S. perplexum, S. pseudocapsicum, S. pseudogracile, S. pumilum, S. rostratum, S. sarrachoides, S. seaforthianum, S. setigeroides, S. sisymbriifolium, S. stoloniferum, S. tampicense, S. tenuipes, S. torvum, S. triflorum, S. triquetrum, S. umbelliferum, S. viarum, S. wallacei
Synonyms S. dulcamara var. villosissimum
Name authority Linnaeus: Sp. Pl. 1: 185. (1753) Dunal in J. Lamarck et al.: Encycl., suppl. 3: 750. (1814)
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