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bitter nightshade, bittersweet, bittersweet nightshade, blue bindweed, climbing nightshade, European bittersweet, felonwort

Douglas' nightshade, greenspot nightshade

Habit Herbs or shrubs, perennial, erect, unarmed, to 1.5(–3) m, sparsely to moderately pubescent, hairs white, curved, unbranched, 0.5–1 mm, eglandular.
Leaves

petiolate;

petiole 0.5–5 cm;

blade simple, elliptic or ovate to cordate, 2.5–12 × 1.2–9 cm, margins entire to deeply pinnatifid and usually 3-lobed near base, lobe margins entire, base truncate to cordate.

petiolate;

petiole 0.5–3(–7) cm;

blade simple, ovate, 1–5(–9) × 0.5–3(–6) cm, margins entire to coarsely and irregularly toothed, base truncate to acute and decurrent.

Inflorescences

terminal or lateral, extra-axillary, much-branched, 7–40-flowered, (1–)4–15 cm.

extra-axillary or leaf-opposed, unbranched, racemelike, 2–7(–14)-flowered, 2–4 cm.

Pedicels

inserted into a small sleeve on the inflorescence axis, 0.6–1.2 cm in flower and fruit.

0.5–1 cm in flower and fruit, nodding or deflexed downward in fruit.

Flowers

radially symmetric;

calyx not accrescent, unarmed, 1–2 mm, glabrous to densely pubescent, lobes triangular, shallow;

corolla purple (rarely white), with green and white shiny spots at base of each lobe, deeply stellate, 1.5–2 cm diam., without interpetalar tissue;

stamens equal;

anthers oblong, slightly tapered, 4.5–6 mm, dehiscent by terminal pores that often open into longitudinal slits;

ovary glabrous.

radially symmetric;

calyx not accrescent, unarmed, 2–3 mm, sparsely pubescent, lobes deltate;

corolla white with yellow-green to brownish central star, stellate, 1–2 cm diam., without interpetalar tissue;

stamens equal;

anthers ellipsoidal and slightly tapered towards the tips, (2.5–)3–4.5 mm, dehiscent by terminal pores that open into longitudinal slits;

ovary glabrous.

Berries

bright shiny red, globose to ellipsoidal, 0.5–1.5 × 0.5–1 cm, glabrous, without sclerotic granules.

dull purplish black, globose, 0.5–1 cm diam., glabrous, with (2–)6–8 sclerotic granules per fruit.

Seeds

pale yellow or tan, flattened, 2–3 mm diam., minutely pitted.

pale yellow to tan, flattened, 1.5–2 × 1–1.5 mm, finely reticulate.

Vines

, climbing or scrambling, herbaceous or woody, unarmed, to 8–10 m, sparsely to densely pubescent, hairs unbranched and/or dendritic, rarely glabrous.

2n

= 24.

= 24.

Solanum dulcamara

Solanum douglasii

Phenology Flowering May–Nov. Flowering Mar–Nov (nearly year-round in Calif.).
Habitat Weedy, in a wide variety of habitats, often associated with water. Dry shrubland, woodland, rocky slopes, stream banks, canyons.
Elevation 0–2000 m. (0–6600 ft.) 0–2500 m. (0–8200 ft.)
Distribution
from FNA
AL; AZ; CA; CO; CT; DC; FL; GA; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; RI; SD; TN; UT; VA; VT; WA; WI; WV; WY; BC; MB; NB; NF; NS; ON; PE; QC; SK; Eurasia
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NM; TX; Mexico; West Indies (Guadeloupe); Central America (El Salvador, Guatemala, Honduras, Nicaragua)
[WildflowerSearch map]
[BONAP county map]
Discussion

Solanum dulcamara is widely distributed across Eurasia and boreal North America. The North American populations are thought to be introductions, but it is possible that the species has a truly circumboreal distribution. A white-flowered form has been recognized by some authors as S. dulcamara forma albiflorum House and an especially pubescent form as S. dulcamara var. villosissimum, but variation in a number of morphological features is continuous across the range of the species and these and other variants are not recognized in the latest monograph of the group (S. Knapp 2013).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Solanum douglasii is most commonly found west of the Rocky Mountains. It can be difficult to distinguish from S. nigrescens, but its longer, slightly tapered anthers on very short (relative to anther length) filaments is a good distinguishing character for plants in flower. The buds of S. douglasii are more pointed than those of S. nigrescens.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 14. FNA vol. 14.
Parent taxa Solanaceae > Solanum Solanaceae > Solanum
Sibling taxa
S. americanum, S. aviculare, S. bahamense, S. campechiense, S. capsicoides, S. carolinense, S. chenopodioides, S. citrullifolium, S. cordicitum, S. davisense, S. deflexum, S. dimidiatum, S. diphyllum, S. donianum, S. douglasii, S. elaeagnifolium, S. emulans, S. erianthum, S. furcatum, S. hindsianum, S. interius, S. jamaicense, S. jamesii, S. laciniatum, S. lanceolatum, S. lumholtzianum, S. lycopersicum, S. marginatum, S. mauritianum, S. nigrescens, S. nigrum, S. nitidibaccatum, S. novomexicanum, S. perplexum, S. pseudocapsicum, S. pseudogracile, S. pumilum, S. rostratum, S. sarrachoides, S. seaforthianum, S. setigeroides, S. sisymbriifolium, S. stoloniferum, S. tampicense, S. tenuipes, S. torvum, S. triflorum, S. triquetrum, S. umbelliferum, S. viarum, S. wallacei
S. americanum, S. aviculare, S. bahamense, S. campechiense, S. capsicoides, S. carolinense, S. chenopodioides, S. citrullifolium, S. cordicitum, S. davisense, S. deflexum, S. dimidiatum, S. diphyllum, S. donianum, S. dulcamara, S. elaeagnifolium, S. emulans, S. erianthum, S. furcatum, S. hindsianum, S. interius, S. jamaicense, S. jamesii, S. laciniatum, S. lanceolatum, S. lumholtzianum, S. lycopersicum, S. marginatum, S. mauritianum, S. nigrescens, S. nigrum, S. nitidibaccatum, S. novomexicanum, S. perplexum, S. pseudocapsicum, S. pseudogracile, S. pumilum, S. rostratum, S. sarrachoides, S. seaforthianum, S. setigeroides, S. sisymbriifolium, S. stoloniferum, S. tampicense, S. tenuipes, S. torvum, S. triflorum, S. triquetrum, S. umbelliferum, S. viarum, S. wallacei
Synonyms S. dulcamara var. villosissimum S. arizonicum
Name authority Linnaeus: Sp. Pl. 1: 185. (1753) Dunal in A. P. de Candolle and A. L. P. P. de Candolle: Prodr. 13(1): 48. (1852)
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