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bitter nightshade, bittersweet, bittersweet nightshade, blue bindweed, climbing nightshade, European bittersweet, felonwort

velvety nightshade, whitetip nightshade

Habit Herbs or shrubs, annual to short-lived perennial, erect or somewhat sprawling, unarmed, to 1 m, glabrescent to densely pubescent, hairs unbranched, to 1 mm, eglandular.
Leaves

petiolate;

petiole 0.5–5 cm;

blade simple, elliptic or ovate to cordate, 2.5–12 × 1.2–9 cm, margins entire to deeply pinnatifid and usually 3-lobed near base, lobe margins entire, base truncate to cordate.

petiolate;

petiole 1–3 cm;

blade simple, narrowly ovate to elliptic, 1.5–5(–7) × 0.5–3.5 cm, margins entire or sinuate, base cuneate to decurrent.

Inflorescences

terminal or lateral, extra-axillary, much-branched, 7–40-flowered, (1–)4–15 cm.

extra-axillary or leaf-opposed, unbranched or rarely forked, umbel-like, 3–7(–10)-flowered, 1–3(–4) cm, fruiting peduncles sharply reflexed from base.

Pedicels

inserted into a small sleeve on the inflorescence axis, 0.6–1.2 cm in flower and fruit.

0.5–1 cm in flower and fruit, reflexed downward in fruit.

Flowers

radially symmetric;

calyx not accrescent, unarmed, 1–2 mm, glabrous to densely pubescent, lobes triangular, shallow;

corolla purple (rarely white), with green and white shiny spots at base of each lobe, deeply stellate, 1.5–2 cm diam., without interpetalar tissue;

stamens equal;

anthers oblong, slightly tapered, 4.5–6 mm, dehiscent by terminal pores that often open into longitudinal slits;

ovary glabrous.

radially symmetric;

calyx not accrescent, unarmed, 2–3.5 mm, sparsely pubescent, lobes deltate, appressed in fruit;

corolla white or purplish, with greenish, yellowish, or brown central star, stellate, 0.8–1.5 cm diam., with sparse interpetalar tissue;

stamens equal;

anthers ellipsoidal, 2–3 mm, dehiscent by terminal pores that open into longitudinal slits;

ovary glabrous.

Berries

bright shiny red, globose to ellipsoidal, 0.5–1.5 × 0.5–1 cm, glabrous, without sclerotic granules.

dull purplish black, globose, 0.5–1 cm diam., glabrous, without sclerotic granules.

Seeds

pale yellow or tan, flattened, 2–3 mm diam., minutely pitted.

pale yellow, flattened, 1–1.5 × 1–1.5 mm, minutely pitted.

Vines

, climbing or scrambling, herbaceous or woody, unarmed, to 8–10 m, sparsely to densely pubescent, hairs unbranched and/or dendritic, rarely glabrous.

2n

= 24.

= 24.

Solanum dulcamara

Solanum chenopodioides

Phenology Flowering May–Nov. Flowering May–Oct (year-round in Fla.).
Habitat Weedy, in a wide variety of habitats, often associated with water. Sandy soil, disturbed areas.
Elevation 0–2000 m. (0–6600 ft.) 0–2000 m. (0–6600 ft.)
Distribution
from FNA
AL; AZ; CA; CO; CT; DC; FL; GA; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; RI; SD; TN; UT; VA; VT; WA; WI; WV; WY; BC; MB; NB; NF; NS; ON; PE; QC; SK; Eurasia
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; FL; GA; MD; MO; NC; WI; South America (Argentina, Brazil, Paraguay, Peru, Uruguay) [Introduced in North America; introduced also in Europe, Africa, Pacific Islands (New Zealand), Australia]
[WildflowerSearch map]
Discussion

Solanum dulcamara is widely distributed across Eurasia and boreal North America. The North American populations are thought to be introductions, but it is possible that the species has a truly circumboreal distribution. A white-flowered form has been recognized by some authors as S. dulcamara forma albiflorum House and an especially pubescent form as S. dulcamara var. villosissimum, but variation in a number of morphological features is continuous across the range of the species and these and other variants are not recognized in the latest monograph of the group (S. Knapp 2013).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Solanum chenopodioides has been introduced sporadically and is occasionally adventive in North America. It is distinctive in having the fruiting peduncles strongly reflexed downward, but is otherwise difficult to distinguish from S. pseudogracile, with which it may be conspecific.

The illegitimate superfluous name Solanum gracile Dunal has often been used for S. chenopodioides (for example, J. K. Small 1913; A. E. Radford et al. 1968). W. G. D’Arcy (1974) included S. gracile (and its replacement name S. ottonis) in the synonymy of S. nigrescens but the taxa are distinct.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 14. FNA vol. 14.
Parent taxa Solanaceae > Solanum Solanaceae > Solanum
Sibling taxa
S. americanum, S. aviculare, S. bahamense, S. campechiense, S. capsicoides, S. carolinense, S. chenopodioides, S. citrullifolium, S. cordicitum, S. davisense, S. deflexum, S. dimidiatum, S. diphyllum, S. donianum, S. douglasii, S. elaeagnifolium, S. emulans, S. erianthum, S. furcatum, S. hindsianum, S. interius, S. jamaicense, S. jamesii, S. laciniatum, S. lanceolatum, S. lumholtzianum, S. lycopersicum, S. marginatum, S. mauritianum, S. nigrescens, S. nigrum, S. nitidibaccatum, S. novomexicanum, S. perplexum, S. pseudocapsicum, S. pseudogracile, S. pumilum, S. rostratum, S. sarrachoides, S. seaforthianum, S. setigeroides, S. sisymbriifolium, S. stoloniferum, S. tampicense, S. tenuipes, S. torvum, S. triflorum, S. triquetrum, S. umbelliferum, S. viarum, S. wallacei
S. americanum, S. aviculare, S. bahamense, S. campechiense, S. capsicoides, S. carolinense, S. citrullifolium, S. cordicitum, S. davisense, S. deflexum, S. dimidiatum, S. diphyllum, S. donianum, S. douglasii, S. dulcamara, S. elaeagnifolium, S. emulans, S. erianthum, S. furcatum, S. hindsianum, S. interius, S. jamaicense, S. jamesii, S. laciniatum, S. lanceolatum, S. lumholtzianum, S. lycopersicum, S. marginatum, S. mauritianum, S. nigrescens, S. nigrum, S. nitidibaccatum, S. novomexicanum, S. perplexum, S. pseudocapsicum, S. pseudogracile, S. pumilum, S. rostratum, S. sarrachoides, S. seaforthianum, S. setigeroides, S. sisymbriifolium, S. stoloniferum, S. tampicense, S. tenuipes, S. torvum, S. triflorum, S. triquetrum, S. umbelliferum, S. viarum, S. wallacei
Synonyms S. dulcamara var. villosissimum S. americanum var. baylisii, S. ottonis
Name authority Linnaeus: Sp. Pl. 1: 185. (1753) Lamarck in J. Lamarck and J. Poiret: Tabl. Encycl. 2: 18. (1794)
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