The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
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bitter nightshade, bittersweet, bittersweet nightshade, blue bindweed, climbing nightshade, European bittersweet, felonwort

apple of sodom, bull nettle, Carolina horse-nettle, Carolina nightshade, devil's tomato, horse nettle

Habit Herbs or shrubs, perennial, erect, sparsely to moderately armed, to 1.2 m, prickles cream to yellowish, straight, to 6 mm, sparsely to densely pubescent, hairs white to cream, sessile to short-stalked, stellate, 4(–8)-rayed, central ray 1–3(–5)-celled and to 3 mm, longer than lateral rays.
Leaves

petiolate;

petiole 0.5–5 cm;

blade simple, elliptic or ovate to cordate, 2.5–12 × 1.2–9 cm, margins entire to deeply pinnatifid and usually 3-lobed near base, lobe margins entire, base truncate to cordate.

petiolate;

petiole 0.4–4 cm;

blade simple, ovate, lanceolate, or elliptic, 2–15 × 2–10 cm, margins subentire, sinuate, or lobed with 1–4 lobes per side, sometimes very deeply lobed almost to midrib, lobe margins entire to coarsely lobed, base cuneate.

Inflorescences

terminal or lateral, extra-axillary, much-branched, 7–40-flowered, (1–)4–15 cm.

extra-axillary, unbranched or rarely forked, 2–12-flowered, 2–9 cm.

Pedicels

inserted into a small sleeve on the inflorescence axis, 0.6–1.2 cm in flower and fruit.

0.5–1 cm in flower, 1.2–1.8 cm and curved downward in fruit.

Flowers

radially symmetric;

calyx not accrescent, unarmed, 1–2 mm, glabrous to densely pubescent, lobes triangular, shallow;

corolla purple (rarely white), with green and white shiny spots at base of each lobe, deeply stellate, 1.5–2 cm diam., without interpetalar tissue;

stamens equal;

anthers oblong, slightly tapered, 4.5–6 mm, dehiscent by terminal pores that often open into longitudinal slits;

ovary glabrous.

radially symmetric;

calyx somewhat accrescent, unarmed or with sparse prickles, 5–8 mm, moderately stellate-pubescent, lobes lanceolate to elliptic-acuminate;

corolla white to pale blue or violet, stellate to stellate-pentagonal or rotate-stellate, 2–3 cm diam., with interpetalar tissue at margins and bases of lobes;

stamens equal;

anthers narrow and tapered, 4.5–6.5 mm, dehiscent by terminal pores;

ovary glabrous or sparsely to moderately glandular-puberulent, rarely moderately pubescent, hairs white, stellate or simple.

Berries

bright shiny red, globose to ellipsoidal, 0.5–1.5 × 0.5–1 cm, glabrous, without sclerotic granules.

light green with darker green mottling or pale greenish white, turning bright yellow, subglobose to depressed-globose, 1–2 × 1–1.8 cm, glabrous, without sclerotic granules.

Seeds

pale yellow or tan, flattened, 2–3 mm diam., minutely pitted.

yellow, flattened, 1.7–2.4 × 1.6–1.8 mm, minutely pitted.

Vines

, climbing or scrambling, herbaceous or woody, unarmed, to 8–10 m, sparsely to densely pubescent, hairs unbranched and/or dendritic, rarely glabrous.

2n

= 24.

= 24.

Solanum dulcamara

Solanum carolinense

Phenology Flowering May–Nov.
Habitat Weedy, in a wide variety of habitats, often associated with water.
Elevation 0–2000 m. (0–6600 ft.)
Distribution
from FNA
AL; AZ; CA; CO; CT; DC; FL; GA; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; RI; SD; TN; UT; VA; VT; WA; WI; WV; WY; BC; MB; NB; NF; NS; ON; PE; QC; SK; Eurasia
[WildflowerSearch map]
[BONAP county map]
from USDA
North America [Introduced in Europe, Asia, Pacific Islands (New Zealand), Australia]
[WildflowerSearch map]
[BONAP county map]
Discussion

Solanum dulcamara is widely distributed across Eurasia and boreal North America. The North American populations are thought to be introductions, but it is possible that the species has a truly circumboreal distribution. A white-flowered form has been recognized by some authors as S. dulcamara forma albiflorum House and an especially pubescent form as S. dulcamara var. villosissimum, but variation in a number of morphological features is continuous across the range of the species and these and other variants are not recognized in the latest monograph of the group (S. Knapp 2013).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf margins subentire, sinuate, or lobed, sinuses of lobes, when present, reaching less than 1/2 distance to midvein; apex of leaf lobes subacute to acute, sometimes rounded.
var. carolinense
1. Leaf margins deeply lobed, sinuses of lobes reaching more than 1/2 distance to midvein or almost to midvein; apex of leaf lobes typically rounded.
var. floridanum
Source FNA vol. 14. FNA vol. 14.
Parent taxa Solanaceae > Solanum Solanaceae > Solanum
Sibling taxa
S. americanum, S. aviculare, S. bahamense, S. campechiense, S. capsicoides, S. carolinense, S. chenopodioides, S. citrullifolium, S. cordicitum, S. davisense, S. deflexum, S. dimidiatum, S. diphyllum, S. donianum, S. douglasii, S. elaeagnifolium, S. emulans, S. erianthum, S. furcatum, S. hindsianum, S. interius, S. jamaicense, S. jamesii, S. laciniatum, S. lanceolatum, S. lumholtzianum, S. lycopersicum, S. marginatum, S. mauritianum, S. nigrescens, S. nigrum, S. nitidibaccatum, S. novomexicanum, S. perplexum, S. pseudocapsicum, S. pseudogracile, S. pumilum, S. rostratum, S. sarrachoides, S. seaforthianum, S. setigeroides, S. sisymbriifolium, S. stoloniferum, S. tampicense, S. tenuipes, S. torvum, S. triflorum, S. triquetrum, S. umbelliferum, S. viarum, S. wallacei
S. americanum, S. aviculare, S. bahamense, S. campechiense, S. capsicoides, S. chenopodioides, S. citrullifolium, S. cordicitum, S. davisense, S. deflexum, S. dimidiatum, S. diphyllum, S. donianum, S. douglasii, S. dulcamara, S. elaeagnifolium, S. emulans, S. erianthum, S. furcatum, S. hindsianum, S. interius, S. jamaicense, S. jamesii, S. laciniatum, S. lanceolatum, S. lumholtzianum, S. lycopersicum, S. marginatum, S. mauritianum, S. nigrescens, S. nigrum, S. nitidibaccatum, S. novomexicanum, S. perplexum, S. pseudocapsicum, S. pseudogracile, S. pumilum, S. rostratum, S. sarrachoides, S. seaforthianum, S. setigeroides, S. sisymbriifolium, S. stoloniferum, S. tampicense, S. tenuipes, S. torvum, S. triflorum, S. triquetrum, S. umbelliferum, S. viarum, S. wallacei
Subordinate taxa
S. carolinense var. carolinense, S. carolinense var. floridanum
Synonyms S. dulcamara var. villosissimum
Name authority Linnaeus: Sp. Pl. 1: 185. (1753) Linnaeus: Sp. Pl. 1: 187. (1753)
Web links