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bitter nightshade, bittersweet, bittersweet nightshade, blue bindweed, climbing nightshade, European bittersweet, felonwort

American black nightshade, American black or common or West Indian nightshade, glossy nightshade, little white nightshade, small-flower nightshade, white nightshade

Habit Herbs to subshrubs, annual to perennial, erect, unarmed, to 1.5 m, glabrate to moderately pubescent, hairs whitish, unbranched, to 1 mm, eglandular.
Leaves

petiolate;

petiole 0.5–5 cm;

blade simple, elliptic or ovate to cordate, 2.5–12 × 1.2–9 cm, margins entire to deeply pinnatifid and usually 3-lobed near base, lobe margins entire, base truncate to cordate.

petiolate;

petiole 1–4 cm;

blade simple, ovate to ovate-elliptic, 2–10.5 × 1–4.5 cm, margins entire or shallowly sinuate-dentate, base decurrent.

Inflorescences

terminal or lateral, extra-axillary, much-branched, 7–40-flowered, (1–)4–15 cm.

extra-axillary or leaf-opposed, unbranched, usually umbel-like, 3–10-flowered, 0.5–3 cm.

Pedicels

inserted into a small sleeve on the inflorescence axis, 0.6–1.2 cm in flower and fruit.

erect or spreading, 0.3–1 cm in flower, to 1.5 cm in fruit.

Flowers

radially symmetric;

calyx not accrescent, unarmed, 1–2 mm, glabrous to densely pubescent, lobes triangular, shallow;

corolla purple (rarely white), with green and white shiny spots at base of each lobe, deeply stellate, 1.5–2 cm diam., without interpetalar tissue;

stamens equal;

anthers oblong, slightly tapered, 4.5–6 mm, dehiscent by terminal pores that often open into longitudinal slits;

ovary glabrous.

radially symmetric;

calyx somewhat accrescent, unarmed, 1–3 mm, lobes deltate, strongly reflexed in fruit;

corolla white, sometimes with yellowish central star, stellate, 0.4–0.8 cm diam., without interpetalar tissue;

stamens equal;

anthers ellipsoidal, 0.7–1.5 mm, dehiscent by terminal pores that open into longitudinal slits;

ovary glabrous.

Berries

bright shiny red, globose to ellipsoidal, 0.5–1.5 × 0.5–1 cm, glabrous, without sclerotic granules.

shiny purplish black, globose, 0.5–1 cm diam., glabrous, with (0–)2–4(–6) sclerotic granules, usually 2–4 larger and 2 smaller.

Seeds

pale yellow or tan, flattened, 2–3 mm diam., minutely pitted.

pale yellow to brown, 1–1.5 × 0.5–1.5 mm, minutely pitted.

Vines

, climbing or scrambling, herbaceous or woody, unarmed, to 8–10 m, sparsely to densely pubescent, hairs unbranched and/or dendritic, rarely glabrous.

2n

= 24.

= 24.

Solanum dulcamara

Solanum americanum

Phenology Flowering May–Nov. Flowering May–Nov (year-round in Fla.).
Habitat Weedy, in a wide variety of habitats, often associated with water. Weedy habitats, secondary forest, disturbed areas.
Elevation 0–2000 m. (0–6600 ft.) 0–1000(–2000) m. (0–3300(–6600) ft.)
Distribution
from FNA
AL; AZ; CA; CO; CT; DC; FL; GA; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; RI; SD; TN; UT; VA; VT; WA; WI; WV; WY; BC; MB; NB; NF; NS; ON; PE; QC; SK; Eurasia
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AZ; CA; FL; GA; LA; MO; MS; OR; SC; TX; UT; WA; BC; Mexico; Central America; South America; West Indies [Introduced in Europe, Asia, Africa, Pacific Islands, Australia]
[WildflowerSearch map]
[BONAP county map]
Discussion

Solanum dulcamara is widely distributed across Eurasia and boreal North America. The North American populations are thought to be introductions, but it is possible that the species has a truly circumboreal distribution. A white-flowered form has been recognized by some authors as S. dulcamara forma albiflorum House and an especially pubescent form as S. dulcamara var. villosissimum, but variation in a number of morphological features is continuous across the range of the species and these and other variants are not recognized in the latest monograph of the group (S. Knapp 2013).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Solanum americanum is a morphologically variable and globally distributed weedy species. It has often been confused with other species in the black nightshade group such as S. emulans, S. nigrescens, S. nigrum, and S. pseudogracile and has often been referred to as S. nodiflorum in floristic treatments. It is distinguished by its very short anthers and shiny black mature fruits with strongly reflexed calyx lobes and usually two to four (rarely none or as many as six) sclerotic granules per fruit. Leaf shape and pubescence can vary considerably throughout its range, and there are some suspected cases of introgression with other species such as S. nigrescens or S. pseudogracile. The name S. nigrum has been misapplied to S. americanum (for example, A. E. Radford et al. 1968).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 14. FNA vol. 14.
Parent taxa Solanaceae > Solanum Solanaceae > Solanum
Sibling taxa
S. americanum, S. aviculare, S. bahamense, S. campechiense, S. capsicoides, S. carolinense, S. chenopodioides, S. citrullifolium, S. cordicitum, S. davisense, S. deflexum, S. dimidiatum, S. diphyllum, S. donianum, S. douglasii, S. elaeagnifolium, S. emulans, S. erianthum, S. furcatum, S. hindsianum, S. interius, S. jamaicense, S. jamesii, S. laciniatum, S. lanceolatum, S. lumholtzianum, S. lycopersicum, S. marginatum, S. mauritianum, S. nigrescens, S. nigrum, S. nitidibaccatum, S. novomexicanum, S. perplexum, S. pseudocapsicum, S. pseudogracile, S. pumilum, S. rostratum, S. sarrachoides, S. seaforthianum, S. setigeroides, S. sisymbriifolium, S. stoloniferum, S. tampicense, S. tenuipes, S. torvum, S. triflorum, S. triquetrum, S. umbelliferum, S. viarum, S. wallacei
S. aviculare, S. bahamense, S. campechiense, S. capsicoides, S. carolinense, S. chenopodioides, S. citrullifolium, S. cordicitum, S. davisense, S. deflexum, S. dimidiatum, S. diphyllum, S. donianum, S. douglasii, S. dulcamara, S. elaeagnifolium, S. emulans, S. erianthum, S. furcatum, S. hindsianum, S. interius, S. jamaicense, S. jamesii, S. laciniatum, S. lanceolatum, S. lumholtzianum, S. lycopersicum, S. marginatum, S. mauritianum, S. nigrescens, S. nigrum, S. nitidibaccatum, S. novomexicanum, S. perplexum, S. pseudocapsicum, S. pseudogracile, S. pumilum, S. rostratum, S. sarrachoides, S. seaforthianum, S. setigeroides, S. sisymbriifolium, S. stoloniferum, S. tampicense, S. tenuipes, S. torvum, S. triflorum, S. triquetrum, S. umbelliferum, S. viarum, S. wallacei
Synonyms S. dulcamara var. villosissimum S. nigrum var. americanum, S. nigrum var. nodiflorum, S. nodiflorum, S. ptychanthum
Name authority Linnaeus: Sp. Pl. 1: 185. (1753) Miller: Gard. Dict. ed. 8 Solanum no. 5. (1768)
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