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tall nightshade, velvety nightshade, whitetip nightshade

bitter nightshade, bittersweet, bittersweet nightshade, blue bindweed, climbing nightshade, European bittersweet, felonwort

Habit Herbs or shrubs, annual to short-lived perennial, erect or somewhat sprawling, unarmed, to 1 m, glabrescent to densely pubescent, hairs unbranched, to 1 mm, eglandular.
Leaves

petiolate;

petiole 1–3 cm;

blade simple, narrowly ovate to elliptic, 1.5–5(–7) × 0.5–3.5 cm, margins entire or sinuate, base cuneate to decurrent.

petiolate;

petiole 0.5–5 cm;

blade simple, elliptic or ovate to cordate, 2.5–12 × 1.2–9 cm, margins entire to deeply pinnatifid and usually 3-lobed near base, lobe margins entire, base truncate to cordate.

Inflorescences

extra-axillary or leaf-opposed, unbranched or rarely forked, umbel-like, 3–7(–10)-flowered, 1–3(–4) cm, fruiting peduncles sharply reflexed from base.

terminal or lateral, extra-axillary, much-branched, 7–40-flowered, (1–)4–15 cm.

Pedicels

0.5–1 cm in flower and fruit, reflexed downward in fruit.

inserted into a small sleeve on the inflorescence axis, 0.6–1.2 cm in flower and fruit.

Flowers

radially symmetric;

calyx not accrescent, unarmed, 2–3.5 mm, sparsely pubescent, lobes deltate, appressed in fruit;

corolla white or purplish, with greenish, yellowish, or brown central star, stellate, 0.8–1.5 cm diam., with sparse interpetalar tissue;

stamens equal;

anthers ellipsoidal, 2–3 mm, dehiscent by terminal pores that open into longitudinal slits;

ovary glabrous.

radially symmetric;

calyx not accrescent, unarmed, 1–2 mm, glabrous to densely pubescent, lobes triangular, shallow;

corolla purple (rarely white), with green and white shiny spots at base of each lobe, deeply stellate, 1.5–2 cm diam., without interpetalar tissue;

stamens equal;

anthers oblong, slightly tapered, 4.5–6 mm, dehiscent by terminal pores that often open into longitudinal slits;

ovary glabrous.

Berries

dull purplish black, globose, 0.5–1 cm diam., glabrous, without sclerotic granules.

bright shiny red, globose to ellipsoidal, 0.5–1.5 × 0.5–1 cm, glabrous, without sclerotic granules.

Seeds

pale yellow, flattened, 1–1.5 × 1–1.5 mm, minutely pitted.

pale yellow or tan, flattened, 2–3 mm diam., minutely pitted.

Vines

, climbing or scrambling, herbaceous or woody, unarmed, to 8–10 m, sparsely to densely pubescent, hairs unbranched and/or dendritic, rarely glabrous.

2n

= 24.

= 24.

Solanum chenopodioides

Solanum dulcamara

Phenology Flowering May–Oct (year-round in Fla.). Flowering May–Nov.
Habitat Sandy soil, disturbed areas. Weedy, in a wide variety of habitats, often associated with water.
Elevation 0–2000 m. [0–6600 ft.] 0–2000 m. [0–6600 ft.]
Distribution
from FNA
CA; FL; GA; MD; MO; NC; WI; South America (Argentina, Brazil, Paraguay, Peru, Uruguay) [Introduced in North America; introduced also in Europe, Africa, Pacific Islands (New Zealand), Australia]
[WildflowerSearch map]
from FNA
AL; AZ; CA; CO; CT; DC; FL; GA; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NM; NV; NY; OH; OR; PA; RI; SD; TN; UT; VA; VT; WA; WI; WV; WY; BC; MB; NB; NF; NS; ON; PE; QC; SK; Eurasia
[WildflowerSearch map]
[BONAP county map]
Discussion

Solanum chenopodioides has been introduced sporadically and is occasionally adventive in North America. It is distinctive in having the fruiting peduncles strongly reflexed downward, but is otherwise difficult to distinguish from S. pseudogracile, with which it may be conspecific.

The illegitimate superfluous name Solanum gracile Dunal has often been used for S. chenopodioides (for example, J. K. Small 1913; A. E. Radford et al. 1968). W. G. D’Arcy (1974) included S. gracile (and its replacement name S. ottonis) in the synonymy of S. nigrescens but the taxa are distinct.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Solanum dulcamara is widely distributed across Eurasia and boreal North America. The North American populations are thought to be introductions, but it is possible that the species has a truly circumboreal distribution. A white-flowered form has been recognized by some authors as S. dulcamara forma albiflorum House and an especially pubescent form as S. dulcamara var. villosissimum, but variation in a number of morphological features is continuous across the range of the species and these and other variants are not recognized in the latest monograph of the group (S. Knapp 2013).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 14. Treatment authors: Lynn Bohs1, 1The author wishes to acknowledge co-authorship with David M. Spooner† on S. jamesii and S. stoloniferum and with Sandra Knapp and Tiina Särkinen on the black nightshade species.. FNA vol. 14. Treatment authors: Lynn Bohs1, 1The author wishes to acknowledge co-authorship with David M. Spooner† on S. jamesii and S. stoloniferum and with Sandra Knapp and Tiina Särkinen on the black nightshade species..
Parent taxa Solanaceae > Solanum Solanaceae > Solanum
Sibling taxa
S. americanum, S. aviculare, S. bahamense, S. campechiense, S. capsicoides, S. carolinense, S. citrullifolium, S. cordicitum, S. davisense, S. deflexum, S. dimidiatum, S. diphyllum, S. donianum, S. douglasii, S. dulcamara, S. elaeagnifolium, S. emulans, S. erianthum, S. furcatum, S. hindsianum, S. interius, S. jamaicense, S. jamesii, S. laciniatum, S. lanceolatum, S. lumholtzianum, S. lycopersicum, S. marginatum, S. mauritianum, S. nigrescens, S. nigrum, S. nitidibaccatum, S. novomexicanum, S. perplexum, S. pseudocapsicum, S. pseudogracile, S. pumilum, S. rostratum, S. sarrachoides, S. seaforthianum, S. setigeroides, S. sisymbriifolium, S. stoloniferum, S. tampicense, S. tenuipes, S. torvum, S. triflorum, S. triquetrum, S. umbelliferum, S. viarum, S. wallacei
S. americanum, S. aviculare, S. bahamense, S. campechiense, S. capsicoides, S. carolinense, S. chenopodioides, S. citrullifolium, S. cordicitum, S. davisense, S. deflexum, S. dimidiatum, S. diphyllum, S. donianum, S. douglasii, S. elaeagnifolium, S. emulans, S. erianthum, S. furcatum, S. hindsianum, S. interius, S. jamaicense, S. jamesii, S. laciniatum, S. lanceolatum, S. lumholtzianum, S. lycopersicum, S. marginatum, S. mauritianum, S. nigrescens, S. nigrum, S. nitidibaccatum, S. novomexicanum, S. perplexum, S. pseudocapsicum, S. pseudogracile, S. pumilum, S. rostratum, S. sarrachoides, S. seaforthianum, S. setigeroides, S. sisymbriifolium, S. stoloniferum, S. tampicense, S. tenuipes, S. torvum, S. triflorum, S. triquetrum, S. umbelliferum, S. viarum, S. wallacei
Synonyms S. americanum var. baylisii, S. ottonis S. dulcamara var. villosissimum
Name authority Lamarck in J. Lamarck and J. Poiret: Tabl. Encycl. 2: 18. (1794) Linnaeus: Sp. Pl. 1: 185. (1753)
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