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downy carrionflower

bristly greenbrier, China root, hellfetter

Habit Herbs.
Stems

annual, climbing, branching, to 2.5 m, herbaceous, glabrous;

prickles absent.

perennial, climbing, branching, terete, to 7+ m × 9 mm, woody, glabrous, with prickles proximally, prickles usually absent distally;

prickles blackish, unequal, 3–10+ mm, bristly, flexible.

Leaves

evenly distributed;

petiole ± equaling blade;

tendrils numerous, long, functional;

blade lustrous, dark green abaxially, narrowly to broadly ovate, 8–16 × 3.5–9 cm, often with minute, whitish pubescence abaxially mostly on veins, with transparent trichomes, base cordate, margins entire, convex, apex long-acuminate;

leaves on branches with oblong, smaller blade.

± persistent in southern part of range, deciduous in north;

petiole 1–2 cm;

blade green, drying to light olive-gray, ovate-lanceolate, ovate, or sometimes pandurate, prominently reticulate, thin, 5–13 × 3–10 cm, not glaucous, glabrous (to prickly on major veins abaxially), base rounded to cordate, margins entire apically, minutely serrulate basally, thin, flat, not banded, not lobed, apex pointed.

Umbels

numerous, axillary to leaves, 10–35-flowered, globose;

peduncle equaling or shorter than subtending leaf.

many, axillary to leaves, to 25-flowered, open to dense, spherical;

peduncle often drooping, 1.5–6.5 cm.

Flowers

perianth greenish;

tepals 3.5–5 mm;

anthers shorter than filaments, ovules (1–)2 per locule;

pedicel 0.5–2.2 cm.

perianth green to bronze;

tepals 4–5 mm;

anthers ± equaling to shorter than filaments;

ovule 1 per locule;

pedicel 0.4–1.2 cm.

Berries

black, subglobose, 8–10 mm, not glaucous.

black, globose, 6–10 mm, not glaucous.

Vines

;

rhizomes knotty, short.

2n

= 26.

Smilax pulverulenta

Smilax tamnoides

Phenology Flowering Apr–Jun. Flowering May–Jun.
Habitat Rich woods, thickets, usually in calcareous soils Wet to dry woods, thickets, bottomlands
Elevation 0–900 m (0–3000 ft) 0–400 m (0–1300 ft)
Distribution
from FNA
AR; DC; DE; IL; IN; KY; MD; MN; MO; NC; NE; NJ; NY; PA; TN; VA; WV
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MN; MO; MS; NC; NE; NJ; NY; OH; OK; PA; SC; SD; TN; TX; VA; VT; WI; WV; ON
[WildflowerSearch map]
Discussion

The distribution of Smilax pulverulenta is disjunct. The eastern distribution is centered in the mid-Atlantic United States, while the western distribution is centered in the Ozark Plateau of Missouri.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Smilax tamnoides is marked by its dark, flexible, and unequal bristles and leaves with minutely serrulate margins basally. The far-western S. californica appears to be closely related but lacks the serrulate margins of the leaf bases.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 26, p. 476. FNA vol. 26, p. 477.
Parent taxa Smilacaceae > Smilax Smilacaceae > Smilax
Sibling taxa
S. auriculata, S. biltmoreana, S. bona-nox, S. californica, S. ecirrata, S. glauca, S. havanensis, S. herbacea, S. hugeri, S. illinoensis, S. jamesii, S. lasioneura, S. laurifolia, S. pseudochina, S. pumila, S. rotundifolia, S. smallii, S. tamnoides, S. walteri
S. auriculata, S. biltmoreana, S. bona-nox, S. californica, S. ecirrata, S. glauca, S. havanensis, S. herbacea, S. hugeri, S. illinoensis, S. jamesii, S. lasioneura, S. laurifolia, S. pseudochina, S. pulverulenta, S. pumila, S. rotundifolia, S. smallii, S. walteri
Synonyms S. herbacea var. pulverulenta S. grandifolia, S. hispida, S. hispida var. australis, S. hispida var. montana, S. tamnoides var. hispida
Name authority Michaux: Fl. Bor.-Amer. 2: 238. (1803) Linnaeus: Sp. Pl. 2: 1030. (1753)
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