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downy carrionflower

bullbrier, carrion-flower, common greenbrier, common greenbrier or catbrier, horsebrier, roundleaf greenbrier

Habit Herbs.
Stems

annual, climbing, branching, to 2.5 m, herbaceous, glabrous;

prickles absent.

perennial, climbing, branching, terete to quadrangular, 5–6+ m × 6 mm, woody, glabrous;

prickles green with dark tips, stout, to 12 mm.

Leaves

evenly distributed;

petiole ± equaling blade;

tendrils numerous, long, functional;

blade lustrous, dark green abaxially, narrowly to broadly ovate, 8–16 × 3.5–9 cm, often with minute, whitish pubescence abaxially mostly on veins, with transparent trichomes, base cordate, margins entire, convex, apex long-acuminate;

leaves on branches with oblong, smaller blade.

deciduous to evergreen, ± evenly disposed;

petiole 0.5–1.5 cm;

tendrils numerous;

blade variable, bright green, drying to pale to brownish green, usually ovate to broadly ovate, with 3 (or 5) ± prominent veins, 4–17 × 4–16 cm, lustrous, not glaucous, glabrous abaxially, base cordate to rounded with acute insertion at petiole, margins entire, apex abruptly pointed.

Umbels

numerous, axillary to leaves, 10–35-flowered, globose;

peduncle equaling or shorter than subtending leaf.

numerous, axillary to leaves, 5–12(–20)-flowered, open to dense, hemispherical to spherical;

peduncle to 1.5 cm, longer or shorter than petiole of subtending leaf.

Flowers

perianth greenish;

tepals 3.5–5 mm;

anthers shorter than filaments, ovules (1–)2 per locule;

pedicel 0.5–2.2 cm.

perianth pale yellowish green to bronze;

tepals 3–4 mm;

anthers shorter than to ± equaling filaments;

ovule 1 per locule;

pedicel 0.2–1.5 cm.

Berries

black, subglobose, 8–10 mm, not glaucous.

blue-black to black, globose, 5–8 mm, glaucous.

Vines

;

rhizomes linear.

2n

= 26.

= 32.

Smilax pulverulenta

Smilax rotundifolia

Phenology Flowering Apr–Jun. Flowering Apr–Jun.
Habitat Rich woods, thickets, usually in calcareous soils Dry to moist, sometimes riparian woods, borders, hedgerows, thickets
Elevation 0–900 m (0–3000 ft) 0–200 m (0–700 ft)
Distribution
from FNA
AR; DC; DE; IL; IN; KY; MD; MN; MO; NC; NE; NJ; NY; PA; TN; VA; WV
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DC; DE; FL; GA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MO; MS; NC; NH; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; WV; NS; ON
[WildflowerSearch map]
[BONAP county map]
Discussion

The distribution of Smilax pulverulenta is disjunct. The eastern distribution is centered in the mid-Atlantic United States, while the western distribution is centered in the Ozark Plateau of Missouri.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Normally, the peduncle of Smilax rotundifolia is about the same length as the petiole of the subtending leaf. In exceptional cases, the peduncle may be considerably longer, thereby making this widely distributed species difficult to distinguish from S. bona-nox and S. tamnoides. It lacks the marginal cartilaginous band found on the leaves of the former species and the hispid prickles of the stem of the latter. Specimens of S. tamnoides lacking prickles may be distinguished by their more strongly ridged stems.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 26, p. 476. FNA vol. 26, p. 476.
Parent taxa Smilacaceae > Smilax Smilacaceae > Smilax
Sibling taxa
S. auriculata, S. biltmoreana, S. bona-nox, S. californica, S. ecirrata, S. glauca, S. havanensis, S. herbacea, S. hugeri, S. illinoensis, S. jamesii, S. lasioneura, S. laurifolia, S. pseudochina, S. pumila, S. rotundifolia, S. smallii, S. tamnoides, S. walteri
S. auriculata, S. biltmoreana, S. bona-nox, S. californica, S. ecirrata, S. glauca, S. havanensis, S. herbacea, S. hugeri, S. illinoensis, S. jamesii, S. lasioneura, S. laurifolia, S. pseudochina, S. pulverulenta, S. pumila, S. smallii, S. tamnoides, S. walteri
Synonyms S. herbacea var. pulverulenta S. caduca, S. quadrangularis, S. rotundifolia var. crenulata, S. rotundifolia var. quadrangularis
Name authority Michaux: Fl. Bor.-Amer. 2: 238. (1803) Linnaeus: Sp. Pl. 2: 1030. (1753)
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