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downy carrionflower

California greenbrier, California smilax, Greenbriar, greenbrier

Habit Herbs. Shrubs or vines; rhizomes short, knotty.
Stems

annual, climbing, branching, to 2.5 m, herbaceous, glabrous;

prickles absent.

perennial, climbing or not, to 12 m, woody, glabrous;

prickles sometimes absent distally, bristlelike, 3–11 mm, flexible.

Leaves

evenly distributed;

petiole ± equaling blade;

tendrils numerous, long, functional;

blade lustrous, dark green abaxially, narrowly to broadly ovate, 8–16 × 3.5–9 cm, often with minute, whitish pubescence abaxially mostly on veins, with transparent trichomes, base cordate, margins entire, convex, apex long-acuminate;

leaves on branches with oblong, smaller blade.

evergreen, ± evenly dispersed;

petiole 0.5–2 cm;

blade dull green, drying to dull, ashy green, ovate to broadly ovate, conspicuously veined, 4–11 × 3–8 cm, not glaucous, glabrous, base cordate to subcordate;

margins entire, thin, flat, not banded, never lobed;

apex acute, often apiculate.

Umbels

numerous, axillary to leaves, 10–35-flowered, globose;

peduncle equaling or shorter than subtending leaf.

axillary to distal leaves, (2–)8–13(–19)-flowered;

peduncle 2–5 cm, longer than petiole of subtending leaf.

Flowers

perianth greenish;

tepals 3.5–5 mm;

anthers shorter than filaments, ovules (1–)2 per locule;

pedicel 0.5–2.2 cm.

perianth green;

tepals 3–6 mm;

ovule 1 per locule;

pedicel thin, 1–1.5 cm.

Berries

black, subglobose, 8–10 mm, not glaucous.

black, ovoid, 7–9 mm.

2n

= 26.

Smilax pulverulenta

Smilax californica

Phenology Flowering Apr–Jun. Flowering May–Jun.
Habitat Rich woods, thickets, usually in calcareous soils Yellow pine and mixed evergreen forests, often in thickets along rivers, streams, and springs, partial–full sun
Elevation 0–900 m (0–3000 ft) 250–1200 m (800–3900 ft)
Distribution
from FNA
AR; DC; DE; IL; IN; KY; MD; MN; MO; NC; NE; NJ; NY; PA; TN; VA; WV
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR
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[BONAP county map]
Discussion

The distribution of Smilax pulverulenta is disjunct. The eastern distribution is centered in the mid-Atlantic United States, while the western distribution is centered in the Ozark Plateau of Missouri.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Smilax californica is apparently closely related to the more eastern S. tamnoides. It lacks the minute serrulations characteristic of the latter’s leaves.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 26, p. 476. FNA vol. 26.
Parent taxa Smilacaceae > Smilax Smilacaceae > Smilax
Sibling taxa
S. auriculata, S. biltmoreana, S. bona-nox, S. californica, S. ecirrata, S. glauca, S. havanensis, S. herbacea, S. hugeri, S. illinoensis, S. jamesii, S. lasioneura, S. laurifolia, S. pseudochina, S. pumila, S. rotundifolia, S. smallii, S. tamnoides, S. walteri
S. auriculata, S. biltmoreana, S. bona-nox, S. ecirrata, S. glauca, S. havanensis, S. herbacea, S. hugeri, S. illinoensis, S. jamesii, S. lasioneura, S. laurifolia, S. pseudochina, S. pulverulenta, S. pumila, S. rotundifolia, S. smallii, S. tamnoides, S. walteri
Synonyms S. herbacea var. pulverulenta S. rotundifolia var. californica
Name authority Michaux: Fl. Bor.-Amer. 2: 238. (1803) (A. de Candolle) A. Gray: in S. Watson, Bot. California 2: 186. (1880)
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