Smilax pulverulenta |
Smilax californica |
|
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downy carrionflower |
California greenbrier, California smilax, Greenbriar, greenbrier |
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Habit | Herbs. | Shrubs or vines; rhizomes short, knotty. |
Stems | annual, climbing, branching, to 2.5 m, herbaceous, glabrous; prickles absent. |
perennial, climbing or not, to 12 m, woody, glabrous; prickles sometimes absent distally, bristlelike, 3–11 mm, flexible. |
Leaves | evenly distributed; petiole ± equaling blade; tendrils numerous, long, functional; blade lustrous, dark green abaxially, narrowly to broadly ovate, 8–16 × 3.5–9 cm, often with minute, whitish pubescence abaxially mostly on veins, with transparent trichomes, base cordate, margins entire, convex, apex long-acuminate; leaves on branches with oblong, smaller blade. |
evergreen, ± evenly dispersed; petiole 0.5–2 cm; blade dull green, drying to dull, ashy green, ovate to broadly ovate, conspicuously veined, 4–11 × 3–8 cm, not glaucous, glabrous, base cordate to subcordate; margins entire, thin, flat, not banded, never lobed; apex acute, often apiculate. |
Umbels | numerous, axillary to leaves, 10–35-flowered, globose; peduncle equaling or shorter than subtending leaf. |
axillary to distal leaves, (2–)8–13(–19)-flowered; peduncle 2–5 cm, longer than petiole of subtending leaf. |
Flowers | perianth greenish; tepals 3.5–5 mm; anthers shorter than filaments, ovules (1–)2 per locule; pedicel 0.5–2.2 cm. |
perianth green; tepals 3–6 mm; ovule 1 per locule; pedicel thin, 1–1.5 cm. |
Berries | black, subglobose, 8–10 mm, not glaucous. |
black, ovoid, 7–9 mm. |
2n | = 26. |
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Smilax pulverulenta |
Smilax californica |
|
Phenology | Flowering Apr–Jun. | Flowering May–Jun. |
Habitat | Rich woods, thickets, usually in calcareous soils | Yellow pine and mixed evergreen forests, often in thickets along rivers, streams, and springs, partial–full sun |
Elevation | 0–900 m (0–3000 ft) | 250–1200 m (800–3900 ft) |
Distribution |
AR; DC; DE; IL; IN; KY; MD; MN; MO; NC; NE; NJ; NY; PA; TN; VA; WV
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CA; OR
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Discussion | The distribution of Smilax pulverulenta is disjunct. The eastern distribution is centered in the mid-Atlantic United States, while the western distribution is centered in the Ozark Plateau of Missouri. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Smilax californica is apparently closely related to the more eastern S. tamnoides. It lacks the minute serrulations characteristic of the latter’s leaves. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 26, p. 476. | FNA vol. 26. |
Parent taxa | Smilacaceae > Smilax | Smilacaceae > Smilax |
Sibling taxa | ||
Synonyms | S. herbacea var. pulverulenta | S. rotundifolia var. californica |
Name authority | Michaux: Fl. Bor.-Amer. 2: 238. (1803) | (A. de Candolle) A. Gray: in S. Watson, Bot. California 2: 186. (1880) |
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