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Blue Ridge carrion-flower

California greenbrier, California smilax, Greenbriar, greenbrier

Habit Herbs. Shrubs or vines; rhizomes short, knotty.
Stems

annual, erect to ascending, branched, 2–2.5 m, herbaceous, glabrous;

prickles absent.

perennial, climbing or not, to 12 m, woody, glabrous;

prickles sometimes absent distally, bristlelike, 3–11 mm, flexible.

Leaves

evenly distributed, proximalmost smaller, narrower;

petiole 1.5–9 cm, shorter than blade;

tendrils numerous, long, functional;

blade not lustrous, pale green abaxially, ovate to round, 4–8 × 3–6 cm, not glaucous, pubescent abaxially, with transparent trichomes, base cordate, margins entire, convex, apex acuminate to rounded and cuspidate.

evergreen, ± evenly dispersed;

petiole 0.5–2 cm;

blade dull green, drying to dull, ashy green, ovate to broadly ovate, conspicuously veined, 4–11 × 3–8 cm, not glaucous, glabrous, base cordate to subcordate;

margins entire, thin, flat, not banded, never lobed;

apex acute, often apiculate.

Umbels

many, axillary to leaves, to 35-flowered, dense, globose;

peduncle to 12+ cm, short.

axillary to distal leaves, (2–)8–13(–19)-flowered;

peduncle 2–5 cm, longer than petiole of subtending leaf.

Flowers

perianth greenish;

tepals 35–45 mm;

anthers equaling or shorter than filaments;

ovules (1–)2 per locule;

pedicel 0.5–1.5 cm.

perianth green;

tepals 3–6 mm;

ovule 1 per locule;

pedicel thin, 1–1.5 cm.

Berries

bluish black to black, subglobose, 8–10 mm, glaucous.

black, ovoid, 7–9 mm.

2n

= 26.

Smilax lasioneura

Smilax californica

Phenology Flowering May–Jun. Flowering May–Jun.
Habitat Rich, alluvial woods, thickets, borders Yellow pine and mixed evergreen forests, often in thickets along rivers, streams, and springs, partial–full sun
Elevation 300–700 m (1000–2300 ft) 250–1200 m (800–3900 ft)
Distribution
from FNA
AL; AR; FL; GA; IA; IL; IN; KS; KY; LA; MI; MN; MO; MS; MT; ND; NE; NY; OH; OK; PA; SD; TN; TX; WI; WY; MB; ON; SK
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from FNA
CA; OR
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[BONAP county map]
Discussion

Smilax lasioneura is primarily distributed in the central plains and lower elevations of the Appalachian and Rocky mountains. Plants distributed in the southern humid areas tend to be larger than those of more northern areas.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Smilax californica is apparently closely related to the more eastern S. tamnoides. It lacks the minute serrulations characteristic of the latter’s leaves.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 26, p. 474. FNA vol. 26.
Parent taxa Smilacaceae > Smilax Smilacaceae > Smilax
Sibling taxa
S. auriculata, S. biltmoreana, S. bona-nox, S. californica, S. ecirrata, S. glauca, S. havanensis, S. herbacea, S. hugeri, S. illinoensis, S. jamesii, S. laurifolia, S. pseudochina, S. pulverulenta, S. pumila, S. rotundifolia, S. smallii, S. tamnoides, S. walteri
S. auriculata, S. biltmoreana, S. bona-nox, S. ecirrata, S. glauca, S. havanensis, S. herbacea, S. hugeri, S. illinoensis, S. jamesii, S. lasioneura, S. laurifolia, S. pseudochina, S. pulverulenta, S. pumila, S. rotundifolia, S. smallii, S. tamnoides, S. walteri
Synonyms Coprosmanthus lasioneuron, Nemexia herbacea subsp. melica, Nemexia lasioneura, Nemexia tenuis, S. diversifolia, S. herbacea var. inodora, S. herbacea subsp. lasioneura, S. herbacea var. lasioneura, S. tenuis S. rotundifolia var. californica
Name authority Hooker: Fl. Bor.-Amer. 2: 173, plate 187A. (1840) (A. de Candolle) A. Gray: in S. Watson, Bot. California 2: 186. (1880)
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