Smilax hugeri |
Smilax californica |
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Huger's carrionflower |
California greenbrier, California smilax, Greenbriar, greenbrier |
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Habit | Herbs; rhizomes knotty, slender. | Shrubs or vines; rhizomes short, knotty. |
Stems | annual, erect, 2–5 m, herbaceous, glabrous; prickles absent. |
perennial, climbing or not, to 12 m, woody, glabrous; prickles sometimes absent distally, bristlelike, 3–11 mm, flexible. |
Leaves | few, disposed distally or evenly distributed, subequal; petiole shorter than blade; tendrils absent or rudimentary; blade oblong, oblong-ovate, or sometimes oval, 2.5–12 × 1.5–7.5 cm, not glaucous, puberulent abaxially, base rounded to slightly cordate, margins entire, apex broadly rounded to obtuse or abruptly pointed. |
evergreen, ± evenly dispersed; petiole 0.5–2 cm; blade dull green, drying to dull, ashy green, ovate to broadly ovate, conspicuously veined, 4–11 × 3–8 cm, not glaucous, glabrous, base cordate to subcordate; margins entire, thin, flat, not banded, never lobed; apex acute, often apiculate. |
Umbels | 1–3, proximalmost axillary to bracts, 5–12-flowered. |
axillary to distal leaves, (2–)8–13(–19)-flowered; peduncle 2–5 cm, longer than petiole of subtending leaf. |
Flowers | perianth greenish; tepals 3–3.5 mm; anthers shorter than filaments; ovules (1–)2 per locule. |
perianth green; tepals 3–6 mm; ovule 1 per locule; pedicel thin, 1–1.5 cm. |
Berries | globose, 8–10 mm diam., glaucous. |
black, ovoid, 7–9 mm. |
2n | = 26. |
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Smilax hugeri |
Smilax californica |
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Phenology | Flowering Apr–May. | Flowering May–Jun. |
Habitat | Moist, mixed woods and wooded slopes | Yellow pine and mixed evergreen forests, often in thickets along rivers, streams, and springs, partial–full sun |
Elevation | 250–1200 m (800–3900 ft) | |
Distribution |
AL; FL; GA; NC; SC; TN
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CA; OR
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Discussion | Smilax hugeri is distinguished from S. ecirrhata and S. biltmoreana mainly by leaf morphology. Specimens from Tennessee are not sharply distinct from S. ecirrhata. J. K. Mangaly (1968) cited this as possible evidence of divergence from that species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Smilax californica is apparently closely related to the more eastern S. tamnoides. It lacks the minute serrulations characteristic of the latter’s leaves. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 26, p. 474. | FNA vol. 26. |
Parent taxa | Smilacaceae > Smilax | Smilacaceae > Smilax |
Sibling taxa | ||
Synonyms | Nemexia hugeri, S. ecirrhata var. hugeri | S. rotundifolia var. californica |
Name authority | (Small) J. B. Norton ex Pennell: Bull. Torrey Bot. Club 43: 420. (1916) | (A. de Candolle) A. Gray: in S. Watson, Bot. California 2: 186. (1880) |
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