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earleaf greenbrier, wild-bamboo

California greenbrier, California smilax, Greenbriar, greenbrier

Habit Shrubs or vines; rhizomes short, knotty.
Stems

perennial, climbing, branching zigzag, terete, 5–9 m × 5–8 mm, woody, glabrous;

prickles numerous, sparse or absent distally, flattened, rigid, stout, to 4 mm.

perennial, climbing or not, to 12 m, woody, glabrous;

prickles sometimes absent distally, bristlelike, 3–11 mm, flexible.

Leaves

evergreen;

petiole 0.5–1.2 cm;

blade green abaxially, drying to brownish green, narrowly ovate to ovate-elliptic, with 3 prominent veins, secondary veins obscure to prominent, 4.5–6(–8.5) × 2–3.5 cm, not glaucous, glabrous or minutely pubescent abaxially, base auriculate, pandurate, or rounded, cuneate at insertion of petiole, margins entire, apex acute to abruptly deflexed point.

evergreen, ± evenly dispersed;

petiole 0.5–2 cm;

blade dull green, drying to dull, ashy green, ovate to broadly ovate, conspicuously veined, 4–11 × 3–8 cm, not glaucous, glabrous, base cordate to subcordate;

margins entire, thin, flat, not banded, never lobed;

apex acute, often apiculate.

Umbels

3–8, terminal, axillary to leaves, 5–8(–25)-flowered, loose;

peduncle 0.2–1.5 cm.

axillary to distal leaves, (2–)8–13(–19)-flowered;

peduncle 2–5 cm, longer than petiole of subtending leaf.

Flowers

perianth green;

tepals: staminate 6–8 mm, pistillate 3–4 mm;

ovule 1 per locule;

pedicel 0.2–1 cm.

perianth green;

tepals 3–6 mm;

ovule 1 per locule;

pedicel thin, 1–1.5 cm.

Berries

purplish maroon, purple, or black, ovoid to flattened, 5–7 mm, glaucous.

black, ovoid, 7–9 mm.

Vines

;

rhizomes linear or dense masses of potatolike tubers.

Smilax auriculata

Smilax californica

Phenology Flowering Apr–Jul. Flowering May–Jun.
Habitat Dunes and sandy flatwoods, full sun Yellow pine and mixed evergreen forests, often in thickets along rivers, streams, and springs, partial–full sun
Elevation 0–100 m (0–300 ft) 250–1200 m (800–3900 ft)
Distribution
from FNA
AL; FL; GA; LA; MS; NC; SC; West Indies (Bahamas)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Smilax californica is apparently closely related to the more eastern S. tamnoides. It lacks the minute serrulations characteristic of the latter’s leaves.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 26, p. 471. FNA vol. 26.
Parent taxa Smilacaceae > Smilax Smilacaceae > Smilax
Sibling taxa
S. biltmoreana, S. bona-nox, S. californica, S. ecirrata, S. glauca, S. havanensis, S. herbacea, S. hugeri, S. illinoensis, S. jamesii, S. lasioneura, S. laurifolia, S. pseudochina, S. pulverulenta, S. pumila, S. rotundifolia, S. smallii, S. tamnoides, S. walteri
S. auriculata, S. biltmoreana, S. bona-nox, S. ecirrata, S. glauca, S. havanensis, S. herbacea, S. hugeri, S. illinoensis, S. jamesii, S. lasioneura, S. laurifolia, S. pseudochina, S. pulverulenta, S. pumila, S. rotundifolia, S. smallii, S. tamnoides, S. walteri
Synonyms S. beyrichii, S. lata S. rotundifolia var. californica
Name authority Walter: Fl. Carol., 245. (1788) (A. de Candolle) A. Gray: in S. Watson, Bot. California 2: 186. (1880)
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