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Habit Plants sometimes canescent basally; caudex branched.
Stems

several from base, unbranched, (0.3–)0.5–1.4(–2) dm, trichomes simple, 0.3–1 mm, mixed with smaller, dendritic ones, (sometimes simple ones absent).

Basal leaves

petiole 0.7–3(–4) cm, often ciliate, trichomes simple;

blade usually linear to oblanceolate or spatulate, rarely obovate, (0.5–)0.8–2.2(–3.5) cm × 0.7–5(–7) mm, margins usually entire, rarely apically 3 or 5-toothed or -lobed, apex obtuse to rounded, (surfaces densely tomentose, grayish, trichomes mostly dendritic, mixed with fewer, simple ones, to 1 mm).

Cauline leaves

shortly petiolate;

blade similar to basal, smaller distally, margins usually entire, rarely lobed.

Racemes

elongated in fruit.

Flowers

sepals 2.5–3.5 mm;

petals white or creamy white, suborbicular to obovate, 4–6 × 2–3(–4) mm, narrowed to claw, 1–2(–3) mm, apex rounded;

anthers oblong, 0.5–0.7 mm.

Fruiting pedicels

divaricate to ascending, (forming 30–70˚ angle), proximalmost bracteate, 4–12 mm, pubescent, trichomes simple mixed with smaller, dendritic ones.

Fruits

divaricate to ascending, ellipsoid to oblong, subterete or slightly 4-angled, 6–10 × 1.5–2.5 mm, base and apex cuneate;

valves each with prominent midvein;

ovules 4–8 per ovary;

style 0.1–0.5 mm.

Seeds

1.5–2 × 0.9–1.2 mm.

2n

= 12, 22, 24.

Smelowskia porsildii

Phenology Flowering Jun–Jul.
Habitat Scree, dry gravelly slopes, sandstone shale-scree, fellfields, gravel beaches and benches, slides, alpine ridges, rock crevices and outcrops, sedge tundra, Dryas heath meadows, dry slopes, tundra slopes, conglomerate outcrops, marshes, sedge meadows, windswept sandstone ridges
Elevation 0-1700 m (0-5600 ft)
Distribution
from FNA
AK; e Asia (Russian Far East)
[BONAP county map]
Discussion

As delimited here, Smelowskia porsildii is taken in the broad sense to include plants recognized by W. H. Drury Jr. and R. C. Rollins (1952) and Rollins (1993) as S. calycina var. integrifolia. The latter was raised by Velichkin to species rank, named as S. spathulatifolia because of the earlier-published S. integrifolia Bunge. Leaf shape and pedicel orientation, used by those authors to distinguish the two taxa, vary continuously. The species is also highly variable in the relative occurrence of simple versus dendritic trichomes and leaf margins. Although I. A. Al-Shehbaz and S. I. Warwick (2006) reduced S. spathulatifolia to synonymy of S. porsildii, the latter most likely represents a complex in which S. media might be involved. Conflicting chromosome numbers (e.g., 2n = 12, 18, 22, 24, 32) have been reported for this complex (Warwick and Al-Shehbaz 2006), and detailed cytological and molecular studies are needed to resolve the taxa involved. The complex involves diploid and tetraploid populations based on x = 6; the counts of 2n = 18 and 32 most likely reflect misidentifications.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 675.
Parent taxa Brassicaceae > tribe Smelowskieae > Smelowskia
Sibling taxa
S. americana, S. borealis, S. johnsonii, S. media, S. ovalis, S. pyriformis
Synonyms S. calycina var. porsildii, Hutchinsia calycina var. integrifolia, S. calycina subsp. integrifolia, S. calycina var. integrifolia, S. jurtzevii, S. spathulatifolia
Name authority (W. H. Drury & Rollins) Jurtsev: Novosti Sist. Vyssh. Rast. 6: 309. (1970)
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