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fernleaf false candytuft

Habit Plants sometimes canescent basally; caudex branched. Perennials [annuals]; eglandular.
Stems

several from base, unbranched, (0.2–)0.5–1.4(–1.8) dm, trichomes simple, 0.5–1.2 mm, mixed with smaller, dendritic ones.

Basal leaves

petiole 0.7–3.5 cm, ciliate, trichomes simple;

blade oblanceolate to obovate, or ovate to oblong in outline, (terminal segments linear, oblong, or ovate), 0.5–3 cm × 4–12 mm, (terminal segments 0.2–1.4 cm × 0.5–4 mm), margins usually pinnatifid, rarely apically 3 or 5-lobed, apex obtuse or subacute.

Cauline leaves

shortly petiolate or sessile;

blade similar to basal, smaller distally.

[rarely absent], petiolate, subsessile, or sessile;

blade base not auriculate, margins usually pinnately or palmately lobed, sometimes entire [dentate].

Trichomes

stalked, usually dendritic, stellate, or forked, sometimes mostly simple.

Racemes

elongated in fruit.

usually ebracteate, often elongated in fruit.

Flowers

sepals 2–3 mm;

petals white, suborbicular to obovate, 4–5 × 2–3 mm, narrowed to claw, 1.5–2 mm, apex rounded;

anthers oblong, 0.5–0.6 mm.

actinomorphic;

sepals suberect to spreading [erect], lateral pair not saccate basally;

petals white, pink, or purple [yellow], claw present, distinct;

filaments unappendaged, not winged;

pollen 3-colpate.

Fruiting pedicels

usually spreading to divaricate, rarely divaricate-ascending, (often forming greater than 40˚ angle, straight or upcurved), proximalmost bracteate, 4–12 mm, pubescent, trichomes simple mixed with smaller, dendritic ones.

Fruits

divaricate-ascending, ellipsoid to oblong, 4-angled, 5–11 × 2–3 mm, base and apex cuneate;

valves each with prominent midvein;

ovules 8–12 per ovary;

style 0.1–0.5 mm.

silicles or siliques, dehiscent, unsegmented, 4-angled, angustiseptate, terete, or subterete [latiseptate];

ovules 4–30[–numerous] per ovary;

style distinct [obsolete];

stigma entire [rarely slightly 2-lobed].

Seeds

1.7–2.2 × 0.9–1.1 mm.

uniseriate [biseriate];

cotyledons accumbent or incumbent.

2n

= 12.

Smelowskia media

Brassicaceae tribe Smelowskieae

Phenology Flowering Jun–Jul.
Habitat Talus scree slopes, dry calcareous gravel flats, limestone scree, tundra turf, seepages, bluffs, stony slopes, grassy quartzite slopes, gravelly lake shores
Elevation 400-1500 m (1300-4900 ft)
Distribution
from FNA
AK; NT; YT
[BONAP county map]
North America; Europe; Asia
Discussion

Smelowskia media resembles S. americana in aspects of habit, flowers, and fruits. It differs by having non-appressed (versus appressed) fruits, fruiting pedicels held at wider angles (greater than 40˚ versus less), and somewhat larger seeds (1.7–2.3 × 0.9–1.2 mm versus 1.1–1.9 × 0.6–0.9 mm).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Genus 1, species 25 (7 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 7, p. 674. FNA vol. 7, p. 671.
Parent taxa Brassicaceae > tribe Smelowskieae > Smelowskia Brassicaceae
Sibling taxa
S. americana, S. borealis, S. johnsonii, S. ovalis, S. porsildii, S. pyriformis
Subordinate taxa
Synonyms S. calycina var. media
Name authority (W. H. Drury & Rollins) Velichkin: Bot. Zhurn. (Moscow & Leningrad) 64: 167. (1979) Al-Shehbaz: Pl. Syst. Evol. 259: 111. (2006)
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