Smelowskia johnsonii
Smelowskia borealis
Johnson's false candytuft
false candytuft
several from base, unbranched or branched proximally, 0.4–1.6 dm, trichomes simple, 1–1.5 mm, mixed with smaller, dendritic ones.
several from base, often unbranched or branched proximally, 0.6–3(–3.5) dm, trichomes simple, to 1 mm, mixed with smaller, dendritic ones.
petiole 0.8–1.3 cm, ciliate, trichomes simple;
blade oblanceolate to spatulate, 0.8–1.8 cm × 3–7 mm, (terminal segments 0.2–0.7 cm × 1–4 mm), margins usually entire or apically 3-toothed or -lobed, rarely palmately 3-lobed, (terminal segments linear to ovate), apex obtuse (surfaces densely villous, silvery, trichomes primarily simple, 1–1.8 mm).
petiole 0.5–3.2 cm, ciliate or not;
blade obovate to ovate in outline, (terminal segments linear, oblong, or ovate), 0.7–2.5 cm × 4–14 mm, (terminal segments 0.2–1.1 cm × 1–5 mm), margins palmately (3 or) 5 (or 7)-lobed, (surfaces densely pubescent, trichomes dendritic, often mixed with larger, simple ones, to 1 mm), apex obtuse or subacute.
subsessile;
blade similar to basal, smaller distally, margins entire.
shortly petiolate or sessile;
blade often similar to basal, smaller distally, margins sometimes pinnatifid.
elongated in fruit.
considerably elongated in fruit.
sepals 3–3.5 mm;
petals lavender to purplish, suborbicular to obovate, 4–5 × 3–4 mm, narrowed to claw, ca. 2 mm, apex rounded;
anthers oblong, 0.5–0.7 mm.
sepals (usually persistent), 2.5–3.2 mm;
petals purple to lavender, suborbicular to obovate, 4–5 × 1.5–2 mm, narrowed to claw, 1.5–2.5 mm, apex rounded;
anthers oblong, 0.4–0.5 mm.
ascending, (often forming less than 40˚ angle, straight), proximalmost bracteate, 11–27 mm, pubescent, trichomes primarily simple (to 1.5 mm).
spreading to divaricate-ascending, (secund, often curved), proximalmost bracteate, 6–15(–20) mm, pubescent, trichomes simple mixed with smaller, dendritic ones.
ascending, ellipsoid to obovoid-ellipsoid, subterete, 5–6 × 2–3 mm, base and sometimes apex cuneate;
valves each with obscure midvein;
ovules 4 per ovary;
style 0.2–0.3 mm.
spreading to ascending, usually oblanceolate to spatulate or linear-oblanceolate, rarely oblong-obovate, angustiseptate, (8–)12–28 × 4–8 mm, base cuneate, apex obtuse;
valves each with prominent midvein and distinct lateral veins, (rarely sparsely pubescent);
ovules 10–18 per ovary;
style 0.1–1.1 mm.
2.2–2.7 × ca. 1 mm.
1.5–2.2 × 1–1.3 mm.
= 12.
Smelowskia johnsonii
Smelowskia borealis
Of conservation concern.
Smelowskia johnsonii is known only from the Bering Strait District, Alaska. It is a distinctive species resembling only superficially the villous forms of S. borealis with simple trichomes. From the latter, S. johnsonii is easily distinguished by having subterete (versus angustiseptate) fruits 2–3 (versus 4–8) mm wide, non-secund (versus secund) and straight (versus often curved) fruiting pedicels, 4 (versus 10–18) ovules per ovary, and simple or apically 3 (or 5)-toothed or -lobed (versus palmately (3 or) 5 (or 7)-lobed) basal leaf blades. It is readily distinguished from all species of the genus by having leaves densely silvery pubescent, with primarily simple trichomes 1–1.8 mm.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
W. H. Drury Jr. and R. C. Rollins (1952) and Rollins (1993) divided Smelowskia borealis into four varieties based on the duration of sepals (persistent versus caducous), style length (less than versus greater than 0.5 mm), and density of indumentum. The various states of these three characters are present in all possible combinations, and they are not inherited in the combinations presented by these authors. Some forms of the species are densely villous and resemble S. johnsonii in their leaf indumentum and perhaps merit recognition as an infraspecific taxon (var. villosa). As delimited here, S. borealis is quite variable; more studies are needed to determine whether or not any of the four varieties merit recognition at some rank. Although numerous collections of the species were examined, none of the types was available for preparing this account. The species is easily distinguished from the remaining North American Smelowskia by having secund infructescences and distinctly angustiseptate fruits.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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